Documenti di Didattica
Documenti di Professioni
Documenti di Cultura
Biology
Eighth Edition
Neil Campbell and Jane Reece
Lectures by Chris Romero, updated by Erin Barley with contributions from Joan Sharp
Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
Overview: Masters of Adaptation
1 µm 2 µm 5 µm
(a) Spherical (b) Rod-shaped (c) Spiral
(cocci) (bacilli)
Prokaryotes Cell-Surface Structures
Carbohydrate portion
of lipopolysaccharide
Outer
Peptidoglycan membrane
Cell Cell
wall
layer wall Peptidoglycan
layer
Plasma membrane Plasma membrane
Protein Protein
Gram- Gram-
positive negative
bacteria bacteria
20 µm
(a) Gram-positive: peptidoglycan (b) Gram-negative: crystal violet is easily
traps crystal violet. rinsed away, revealing red dye.
toxic bacteria
• A polysaccharide or protein layer called a
capsule covers many prokaryotes.
• Some prokaryotes have fimbriae (also called
attachment pili), which allow them to stick to
their substrate or other individuals in a colony
• Sex pili are longer than fimbriae and allow
prokaryotes to exchange DNA by conjugation.
Fimbriae
200 nm
Motility / Internal Organization
Flagellum
Filament
50 nm
Cell wall
Hook
Basal apparatus
Plasma
membrane
Bacteria often have Specialized Internal Membranes
0.2 µm 1 µm
Respiratory
membrane
Thylakoid
membranes
1 µm
Reproduction and Endospores Adaptation
Endospore
0.3 µm
Bacteria
EXPERIMENT
Evolve Rapidly
in Response to Daily serial transfer
Environmental 0.1 mL
Conditions (population sample)
RESULTS
1.8
Fitness relative
1.6
to ancestor
1.4
1.2
1.0
A+ B+
A+ B+
Bacteria Donor
Cell
A+
Recombination
A+
A– B–
Bacteria Recipient
Cell
A+ B–
Recombinant cell
Conjugation and Plasmids
Sex pilus 1 µm
Conjugation
F+ cell F+ cell
Mating
bridge
F– cell F+ cell
Bacterial
chromosome
Recombinant
Hfr cell A+ A+ A+ F– bacterium
A+
F factor A– A+
A– A– A+ A–
F– cell
Photosynthetic
cells
Heterocyte
20 µm
Lessons from Molecular Systematics
Eukarya
Domain
Systematics: Eukaryotes
Classification
Korarcheotes
Domain Archaea
Euryarchaeotes
Crenarchaeotes
UNIVERSAL Nanoarchaeotes
ANCESTOR
Proteobacteria
Domain Bacteria
Chlamydias
Spirochetes
Cyanobacteria
Gram-positive
bacteria
Domain: Archaea
Eukarya
Archaea
Bacteria
Extreme Thermophiles
• Methanogens live in swamps and marshes
and produce methane as a waste product.
• Methanogens are strict anaerobes and are
poisoned by O2
• In recent years, genetic prospecting has
revealed many new groups of archaea.
• Some of these may offer clues to the early
evolution of life on Earth.
CHLAMYDIAS SPIROCHETES
2.5 µm
5 µm
Chlamydia (arrows) inside an Leptospira, a spirochete
animal cell (colorized TEM) (colorized TEM)
1 µm
5 µm
Two species of Oscillatoria, Streptomyces, the source of Hundreds of mycoplasmas
filamentous cyanobacteria (LM) many antibiotics (colorized SEM) covering a human fibroblast
cell (colorized SEM)
Cyanobacteria
50 µm
Two species of Oscillatoria,
filamentous cyanobacteria (LM)
Gram-Positive Bacteria
5 µm
• Pathogenic prokaryotes typically cause disease
by releasing exotoxins or endotoxins.
• Exotoxins cause disease even if the
prokaryotes that produce them are not present.
• Endotoxins are released only when bacteria
die and their cell walls break down.
• Many pathogenic bacteria are potential
weapons of bioterrorism.
Fimbriae
Cell wall
Circular chromosome
Capsule
Sex pilus
Internal
organization
Flagella
Mutualism ++ Nitrogen Fixing Bacteria and Legume Plants
You should now be able to: