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• hemoglobin
Nervous System:
Circular muscle
Longitudinal muscle
Digestive System: • Crop = “stomach”;
stores food
• Gizzard = grinds food
• Intestine = nutrient
absorption
• Anus
Excretory System:
• A pair of nephridia per segment
• Includes:
– Nephridia
– Bladder
– Excretory pore
Reproductive System:
• Reproductive –
– Hermaphrodites
– Trochophore larvae
Stomach
Ciliary band
Mouth
Anus
1
5
Annelid development
IV. Classes
A. Class Polychaeta (many bristles) Hermodice
– most numerous # species crunculata
– marine
Class Polychaeta
trophosome
Riftia pachyptila
Polychaete Reproduction
• Dioecious
• Trochophore larvae
B. Class Clitellata
1. Subclass Oligochaeta (few bristles)
• Freshwater, marine & terrestrial
Lumbricus terrestris
Class Oligochaeta
• Defining characteristics
– Cylindrical glandular region
of the body = clitellum
• Second largest class in the
phylum Annelida
• Most spp. are earthworms,
very few are marine
Class Oligochaeta
Oligochaete Reproduction
Oligochaete Development
• For terrestrial oligochaetes, development is
direct without any larval forms
• Other Structures:
• Setae: bristles used for movement
(ventral side)
• Clittelum: enlarged segment;
secretes mucus & contains
reproductive parts
Annelid Taxomony
B. Class Clitellata
2. Subclass Hirudinea
• Fixed # segments (34)
• Setae absent
Hirudo
medicinalis
Class Hirudinea
• Defining characteristics
– Posterior sucker
– Anterior sucker
– No setae
– hermaphrodites
• Predominately freshwater, but do
occur in all seas and moist soil
• Leeches do not burrow or crawl,
lack parapods and setae
• Dorsoventrally compressed (flat)
• Feed on invertebrates, body fluids,
32blood Phylum Annelida
Subclass Hirudinea
Leech Anatomy
34 Phylum Annelida
Blood Sucker
• The salivary glands excrete hirudin which prevents the
blood from clotting
• Leeches were commonly used in the 19th century for
bloodletting
– Recent medical uses are to relieve pressure after vascular
tissue is damaged
35 Phylum Annelida
Leeches
o Parasitic segmented worm
o Most live in fresh water
o Have no bristles for movement
o move with muscular
contractions
o Suckers used for attachment to
host
o Some have suckers on tail also
o http://arkive.org/medicinal-
leech/hirudo-
medicinalis/video-00.html
o Secrete anesthetic during bite and
release anticoagulant into blood
during feeding (to keep blood
flowing)
Dissection
Cross section