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BioenergeticsLecture5summary

Susan.Kaminskyj@usask.ca
Lasttimeintegratingcatabolic
metabolism,reviewcatabolism
Thistime

similaritiesanddifferencesbetweencellular
respirationandphotosynthesis
chlorophyll,photosystemsIIandI(inthat
order)

Butfirst.!

ElectrontransportchainpumpsH+,makingagradient
ATPsynthaseusestheH+gradienttogenerateATP
HigherH+concentration=lowerpH
Intermembranespace

Potentialenergyof
fallingH+
usedtogenerateATP

Potentialenergyof
fallingwater
usedtogrindgrain

F1in
matrix

Mitochondrialmatrix

Foininner
membrane

LowerH+concentration=higherpH

ATPsynthaseusestheH+gradienttogenerateATP

Cellular Respiration

Aseriesofredoxreactions.
RESPIRATION
PHOTOSYNTHESIS

PHOTOSYNTHESIS
RESPIRATION

Respirationvsphotosynthesis
Mitochondria
Allaerobiceukaryotes

Chloroplasts
Plants,algae

OxidationofCHOtoCO2 Energyharvestfromsunlight
GenerationofNADHand GenerationofNADPH
FADH2
ATPsynthesis

ReductionofCO2toCHO

CHO=carbohydrate
Pphotosynthesis
NoteNADH(respiration)vsNADPH(photosynthesis)

Photosynthesisisan
endothermicredoxprocess

Energysource?
Usefulbyproducts?

Sunlight
GlucoseandO2

RedandBlue
lightare
absorbedfrom
theincident
(white)light,
leavingGreen
tobereflected
ortransmitted

Photosynthesisproducesoxygen
Areall
wavelengths
photosynthetically
active?
Spirogyra

Photosynthetic
Active
spectrum

First,lightenergymustbecaptured

Illuminatedchlorophyllfluoresces
(givesofflight)ifcapturedlight
energyisnottransferredto
anotheracceptor

Captureisnotenough!
Inaleaf,the
reactioncentre
transfersthe
capturedenergy
toarelatively
stable
intermediate
chemical

Isolatedchlorophyllfluoresces
ifnewlycapturedenergyisnottransferred

Thereactioncentreistheheartof
thephotosystem

Energytransfer
betweenpigment
moleculestoa
specialcentralpair
ofchlorophylls
(reactioncentre)
andthencetothe
primaryelectron
acceptors

Twopartstothelightreaction

ChemiosmosiscanmakeATP

Twopartstothelightreaction
phaeophytin

ferredoxin

Sourceandfate
ofcarbonand
oxygenin
carbohydrates
formedby
photosynthesis

PhotosystemIImakesATP

PhotosystemImakesNADPH

NADHvsNADPH
NADHcatabolism
NADPHanabolism
(Pphotosynthesis)
Foreach,thereduced
formstores~3times
moreenergythanATP

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