Brailovsky & Perez-Gelabert 2019 Review Coreidae of Hispaniola PDF

Zootaxa 4568 (2): 201–241 ISSN 1175-5326 (print edition)
https://www.mapress.com/j/zt/
Copyright © 2019 Magnolia Press
Article ZOOTAXA
ISSN 1175-5334 (online edition)
https://doi.org/10.11646/zootaxa.4568.2.1
http://zoobank.org/urn:lsid:zoobank.org:pub:3265C7BB-159A-4C23-9167-D479484B6382
A review of the Coreidae of Hispaniola (Hemiptera: Heteroptera),

with description of one new species, new distributional records,
and a key to the subfamilies, tribes, genera and species
HARRY BRAILOVSKY1 & DANIEL E. PEREZ-GELABERT2

1
Instituto de Biología, Departamento de Zoología, Universidad Nacional Autónoma de México, Apdo Postal 70153, Ciudad de Méxi-
co, 04510. México. E-mail: coreidae@ib.unam.mx.
2
Integrated Taxonomic Information System (ITIS) and Department of Entomology, National Museum of Natural History, Smithsonian
Institution, Washington, D. C., USA. E-mail: perezd@si.edu.
Table of content
Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 202
Resumen . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 202
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 203
Material and methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 204
Coreidae of Hispaniola . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 204
Subfamily Meropachyinae. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 205
Tribe Merocorini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 205
Merocoris Perty 1830 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 205
Key to Hispaniola species of Merocoris (Corynocoris) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 205
Merocoris (C.) distinctus Dallas . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Merocoris (C.) typhaeus (Fabricius) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Subfamily Coreinae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Tribe Acanthocerini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Acanthocerus Palisot de Beauvois 1818 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 206
Acanthocerus crucifer Palisot de Beauvois . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 207
Tribe Anisoscelini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 209
Key to Hispaniola genera and species of Anisoscelini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 209
Chondrocera Laporte 1832 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 209
Chondrocera mirabilis Brailovsky & Barrera. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 210
Leptoglossus Guérin-Méneville 1831 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 210
Key to Hispaniola species of Leptoglossus . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 210
Leptoglossus balteatus (Linnaeus) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 211
Leptoglossus concolor (Walker) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 212
Leptoglossus confusus Alayo & Grillo . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 212
Leptoglossus gonagra (Fabricius). . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 213
Mamurius Stål 1862 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 214
Mamurius cubanus Barber & Bruner . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 214
Phthia Stål 1862 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 214
Phthia rubropicta (Westwood) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 215
Phthiacnemia Brailovsky 2009 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 215
Phthiacnemia picta (Drury) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 215
Rhytidophthia Brailovsky 2009 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 216
Rhytidophthia splendida (Valdes) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 216
Tribe Chariesterini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 217
Chariesterus De Laporte 1832 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 217
Chariesternus gracilicornis Stål . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 217
Tribe Hypselonotini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220
Key to Hispaniola Genera of Hypselonotini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220
Althos Kirkaldy 1904. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220
Althos obscurator (Fabricius) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220
Anasa Amyot and Serville 1843 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 221
Accepted by D. Rider: 8 Feb. 2019; published: 20 Mar. 2019 201

Key to Hispaniola Species of Anasa . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 221
Anasa acutangula Stål. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 222
Anasa bellator (Fabricius). . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 222
Anasa pisina Brailovsky . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 223
Anasa scorbutica (Fabricius) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 223
Anasa tristis (De Geer) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 225
Catorhintha Stål 1859 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 225
Key to Hispaniola Species of Catorhintha . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 225
Catorhintha guttula (Fabricius) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 225
Catorhintha selector Stål. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 226
Sethenira Spinola 1837 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 227
Sethenira ferruginea Stål . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 227
Zicca Amyot and Serville 1843 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 227
Key to Hispaniola Species of Zicca . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 228
Zicca gloriosa Brailovsky & Perez-Gelabert sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 228
Zicca rubricator rubricator (Fabricius) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 229
Zicca taeniola (Dallas) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 229
Tribe Spartocerini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231
Key to Hispaniola Genera of Spartocerini . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231
Eubule Stål 1867 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231
Eubule spartocerana Brailovsky . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 231
Spartocera Laporte 1832 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 232
Key to Hispaniola Species of Spartocera . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 232
Spartocera batatas (Fabricius) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 232
Spartocera fusca (Thunberg) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 233
Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 234
Check list of the known subfamilies, tribes, genera and species of Dominican Republic . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 237
Check list of the known subfamiies, tribes, genera and species of Haiti . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 238
Abstract
A faunistic survey of the Hispaniolan bugs of the family Coreidae is presented based on the study of over 1000 specimens
deposited in several entomological collections. The new species Zicca gloriosa sp. nov. is described from the Dominican
Republic. Another 12 species are new records for the island, 11 of them being new records for the Dominican Republic:
Althos obscurator (Fabricius, 1803), Anasa acutangula Stål, 1870, Anasa tristis (De Geer, 1773), Eubule spartocerana
Brailovsky, 1992, Leptoglossus confusus Alayo & Grillo, 1977, Mamurius cubanus Barber & Bruner, 1947, Merocoris
distinctus Dallas, 1852, Merocoris typhaeus (Fabricius, 1798), Phthia rubropicta (Westwood, 1842), Sethenira ferruginea
Stål, 1870, and Zicca rubricator rubricator (Fabricius, 1803). New records for Haiti are Anasa scorbutica (Fabricius,
1798), Catorhintha selector Stål, 1859, Chariesterus gracilicornis Stål, 1870, Rhytidophthia splendida (Valdes, 1910) and
Zicca taeniola (Dallas, 1852). The literature citations presented in Perez-Gelabert (2008) of Leptoglossus cinctus (Her-
rich-Schäffer, 1836) and Leptoglossus stigma (Herbst, 1784) are probably based on erroneous identifications and not
counted as part of the Hispaniolan fauna. The total number of species presently known from Hispaniola is raised to 28.
Key to subfamilies, tribes, genera and species are included, as well as color dorsal habitus.
Resumen
Se presenta un inventario de la fauna de chinches de la familia Coreidae en la Hispaniola basado en el estudio de más de
mil especímenes depositados en varias colecciones entomológicas. Se describe la nueva especie Zicca gloriosa sp. nov.
de la República Dominicana. Además, 12 especies son nuevos registros para la isla, 11 de ellas son nuevos registros para
la República Dominicana: Althos obscurator (Fabricius, 1803), Anasa acutangula Stål, 1870, Anasa tristis (De Geer,
1773), Eubule spartocerana Brailovsky, 1992, Leptoglossus confusus Alayo & Grillo, 1977, Mamurius cubanus Barber
& Bruner, 1947, Merocoris distinctus Dallas, 1852, Merocoris typhaeus (Fabricius, 1798), Phthia rubropicta (Westwood,
1842), Sethenira ferruginea Stål, 1870, y Zicca rubricator rubricator (Fabricius, 1803). Nuevos registros para Haití son:
Anasa scorbutica (Fabricius, 1798), Catorhintha selector Stål, 1859, Chariesterus gracilicornis Stål, 1870, Rhytidophthia
splendida (Valdes, 1910) y Zicca taeniola (Dallas, 1852). Las citas de la literatura presentadas en Perez-Gelabert (2008)
de Leptoglossus cinctus (Herrich-Schäffer, 1836) y Leptoglossus stigma (Herbst, 1784) son probablemente basadas en
identificaciones erróneas y descontadas como parte de la fauna en la Hispaniola. El número total de especies conocidas
202 · Zootaxa 4568 (2) © 2019 Magnolia Press BRAILOVSKY & PEREZ-GELABERT
de la Hispaniola se incrementa a 28. Clave para subfamilias, tribus, géneros y especies son incluidas, así como fotografías
en vista dorsal.
Key words: Coreid bugs, faunal survey, Dominican Republic, Haiti, Greater Antilles, Caribbean
Introduction
Bugs of the family Coreidae are commonly known as squash bugs, leaf-footed bugs or plant-feeding bugs and
usually are phytophagous sap-sucking hemipterans that vary from medium to large in size and feed on a wide
variety of plants. Both nymphs and adults suck sap from foliage, flowers, fruits and seeds, affecting cultivated
plants and fruit production. Occasional reports mention that some individuals imbibe fluids from descomposing
animal carcasses. Although normally not a severe problem, some species have the potential to become serious
pests. Adults and nymphs are capable of exuding a strong-smelling fluid when disturbed (Froeschner 1988 in
Henry & Froeschner 1988); Schuh & Slater 1995; Brailovsky 2011).
The coreids are characterized by having the head usually small, relative to body size; antennae inserted above
a line running through the center of eyes; rostrum with four segments; hind femur sometimes ornamented with
large spines; hemelytral membrane usually with numerous veins; well developed scent glands opening ventrally on
the metathorax of adults; nymphal dorsal abdominal scent glands orifices between terga IV–V and V–VI; and all
abdominal spiracle ventral. The body sizes include the largest living heteroperans, as well as other species that are
delicate or slender. The body color is variable (Levin Mitchell in Schaefer & Panizzi 2000; Schuh & Slater 1995;
Brailovsky 2011)
At present, the known global diversity of Coreidae includes 2,563 valid species classified in 432 genera,
making them the fourth largest family of true bugs (Heteroptera) after the Miridae (>10,000 species), Reduviidae
(~7500 species) and Tingidae (~2,600 species).
Hispaniola is the second largest island of the West Indies (Greater Antilles, Lesser Antilles and Bahamas), with
nearly 78,000 km2 and is located in the Caribbean Basin just under the Tropic of Cancer within 17º 40’ and 19º 56’
North latitude and 68º 20’ and 72º 01’ West longitude (Fig. 1). Two countries are the political tenants of this natural
geographic unit: Haiti in the western one-third and the Dominican Republic on the eastern two-thirds of the
territory. The remarkable physiographic variability of its terrain includes the highest and most extensive mountain
systems in the West Indies. Four cordilleras extend across the island in a northwest – southeast axis with valleys in
between. Some peaks reach upwards of 3000 m, while there are also areas below sea level. This mosaic of
microclimates and ecosystems throughout the island harbors a rich flora and fauna with significant numbers of
endemic species. Two major and previously independent terranes or paleoislands coalesced to form modern
Hispaniola. When brought together, the total biological diversity of this territory increased. At present, many areas
of this insular environment have been significantly altered by agriculture, introduction of exotic plant and animal
species, and urbanization. A total of 5,676 insect species were recorded from Hispaniola by Perez-Gelabert (2008)
in the first checklist ever to produce a quantification of the existing knowledge of its arthropod fauna. Despite the
important advance, the insect fauna of Hispaniola still is the least known of the Greater Antilles.
In the islands of the West Indies, the diversity of Coreidae is relatively low, only including some 60 species in
24 genera classified in the subfamilies Meropachyinae and Coreinae. Seventeen species of Coreidae were included
in the list of Perez-Gelabert (2008), although two are probable misidentifications.
The characterization of West Indian coreids started with some of the classical works of Hemiptera taxonomy in
the 19th century, which included the first descriptions of species from these islands. For example, Palisot de
Beauvois (1805–1822) described one of the most common coreids in Hispaniola, Acanthocerus crucifer in 1818.
The most significant contributions to the taxonomic knowledge of Coreidae in the West Indian area are the reviews
of Ruckes (1955) on Chariesterus and the works of Brailovsky (some of these with collaborators) on Anasa (1985,
2001, 2008), Catorhintha (1987), Sphictyrtus (1989), Althos (1990), Zicca (1992), Chondrocera (1994), Phthia
complex (2009) and Leptoglossus (2014), among other works. Faunistic studies of coreids in the West Indies were
summarized most thoroughly in Cuba by Barber & Bruner (1947); Alayo (1967); Bruner, Scaramuzza & Otero
(1975); Alayo & Grillo (1977) and Grillo (2012). Various works included species from Puerto Rico and other
islands (Barber 1923, 1926; Allen 1969; O’Shea 1980; Halbert & Brambila 2005). The compilation of Wolcott
(1948) for Puerto Rican insects contains the most comprehensive summary of coreids in that island. As for many
A REVIEW OF THE COREIDAE OF HISPANIOLA Zootaxa 4568 (2) © 2019 Magnolia Press · 203
other insect groups, a faunistic study and thorough sampling of coreids in Hispaniola has never been attempted.
Due to the paucity of insect collections from Haiti, that part of the island is even less known. The only existing
summary of the Coreidae fauna of Hispaniola is found within the checklist of Hispaniolan arthropods by Perez-
Gelabert (2008). A more thorough sampling of ecologically unique forest areas is expected to yield still unknown
species.
The Coreidae of the Dominican Republic include two subfamilies, seven tribes, 16 genera, and 28 species, one
of them new to science. The Coreidae of Haiti include one subfamily, four tribes, seven genera and eight species all
but Rhytidophthia splendida also cited from the Dominican Republic. The purpose of this work is to characterize
the extent of diversity of the Hispaniolan fauna of Coreidae based on the study of a large number of specimens.
FIGURE 1. Map of the island of Hispaniola.
Material and methods
The diversity of Coreidae in Hispaniola was assessed through the examination of over 1000 specimens from
multiple collections in the Dominican Republic, United States, Mexico and Europe.
The following acronyms are used for the institutions cited here: Natural History Museum, London, England
(BMNH); California Academy of Sciences (Golden Gate Park), San Francisco, California, USA (CASC);
California State Collection, Department of Food & Agriculture, Sacramento, California, USA (CDFA); Carnegie
Museum of Natural History, Pittsburgh, PA, USA (CMNH); Field Museum of Natural History, Chicago, Illinois,
USA (FMNH); Florida State Collection of Arthropods, Gainesville, Florida, USA (FSCA); Instituto de
Investigaciones Botánicas y Zoológicas, Santo Domingo, República Dominicana (IIBZ); Museo Nacional de
Historia Natural de Santo Domingo, República Dominicana (MNHNSD); Texas A & M University (College
Station), USA (TAMU); University of Georgia, Athens, Georgia, USA (UGAG); University of Minnesota, Insect
Collection, USA (UMSP); Instituto de Biología, Universidad Nacional Autónoma de México (UNAM); National
Museum of Natural History, Smithsonian Institution, Washington, D. C., USA (USNM), and Zoologisches Institut
und Museum, University of Hamburg, Germany (ZIZM). All measurements are given in mm.
Coreidae of Hispaniola
Key to Hispaniola subfamilies, tribes, genera and species of Coreidae* »»
1. Hind tibiae with a distinct tooth or spine at distal ends (may be obscured by setae); hind femora longer than abdomen, narrow
and cylindrical basally, swollen apically; body small, robust, thick, somewhat compressed laterally; scutellum more or less
raised, terminating in a single elevated knob . . . . . . . . . . . . . . . . . . . . Meropachyinae Stål...Merocorini Stål...Merocoris Perty
- Hind tibiae lacking a distinct spine or tooth at distal ends . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Coreinae Leach...2
2. Hind femora spinose or tuberculate, strongly incrassate in males; head not over half as long as pronotum and not extending
past antenniferous tubercles or very slightly so; antenniferous tubercles occupying most of anterior head width, space between
them rarely wider than one tubercle; antenniferous tubercles usually with a spine present; ocellar tubercle large; metathoracic
scent gland with a single fused auricle; parameres membranous . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Acanthocerini Bergroth...Acanthocerus crucifer Palisot de Beauvois
- Not with this combination of characters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
3. Head long, longer than wide, anterior portion always porrect; tylus and juga porrect never deflexed or projecting upward in the
form of a triangular spine; head usually longer than scutellum; hind femora reaching the apex of abdomen; tibiae usually folia-
ceous, expanding both dorsally and ventrally or tibiae slightly or not expanded; all femora ventrally armed, fore femora spines
sometimes minute; buccula never reaching midlength of head . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Anisoscelini Laporte
- Not with this combination of characters . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
4. Neither juga nor tylus strongly deflexed; head not appearing incised anteriorly; antenniferous tubercles not prominently pro-
duced; spiracles equidistant from anterior to posterior borders of middle abdominal segments . . . . . . . Hypselonotini Bergroth
- Juga and tylus strongly and abruptly deflexed; head appearing incised anteriorly; antenniferous tubercles prominently pro-
duced; spiracles situated close to anterior border of middle abdominal segments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
5. Distance between hind coxae equal to distance from coxa to lateral margn; antenniferous tubercles prominent, subcontiguous
above deflexed tylus and jugum; antennal segment III cylindrical, not dilated distally; antenniferous tubercles lacking a distinct
spine . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Spartocerini Amyot & Serville
- Distance between hind coxae much narrower than distance from coxa to lateral margin; antennal segment III expanded; anten-
niferous tubercle usually bearing a distinct spine . . . . . . . . . . . . . . . . . . . . . .Chariesterini Stål...Chariesterus gracilicornis Stål
*Modified from Packauskas key (1994)

»» The species are included in this key only if a single species is recorded for that genus; otherwise in the treatment of each
genus with more than one known species a specific key is added.
Subfamily Meropachyinae
Tribe Merocorini
Discussion. The tribe is represented by a single genus, Merocoris Perty 1833, with two subgenera and eight
species. The subgenus Merocoris (Perty 1833) is characterized by having the antenniferous tubercle unarmed, the
anterolateral borders of the pronotum smooth or with a single tubercle, and the scutellar disk conically raised. It
includes five South American species. The subgenus Corynocoris (Mayr 1866) has the antenniferous tubercles
externally armed with a stout spine, the anterolateral borders of the pronotum with two or more tubercles, and the
scutellar disk almost flat. The genus includes three species distributed in North America, Central America and the
Greater Antilles (Brailovsky and Barrera 2009).
Merocoris Perty 1830
Merocoris (Corynocoris) Mayr 1866
Diagnosis. Body stout, thickly pubescent. Hind femora curved, slender on proximal half, strongly swollen distally
with a series of large ventral spines. Head short, armed with a short sharp spine at base. Pronotum strongly
declivent. Scutellum triangular, short, reaching onto base of claval commissure, with apex elevated as a small
knob. Scutellar disk carinate mesally, irregularly sculptured laterally. Abdominal spiracles circular, tiny, and
located between the anterior and middle third of each sternite.
Key to Hispaniola species of Merocoris (Corynocoris)
1. Head subtriangular prolonged anteriorly between the bases of the antenniferous tubercles; interocular space with short tubercle
near the inner border of each eye; antennal segment I subclavate, tapering toward the base from the middle and finely pubes-
cent . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . M. (C.) typhaeus (Fabricius)
- Head quadrate, not conspicuously prolonged in front of the bases of the antenniferous tubercles; interocular space unarmed;
antennal segment I stout, constricted basally only, and densely pubescent with hairs thick and bristle-like . . . . . . . . . . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . M. (C.) distinctus Dallas
Merocoris (C.) distinctus Dallas

(Fig. 2)
Merocoris distinctus Dallas, 1852: 419.
Discussion. Under the known Hispaniola species of Merocoris, this species is distinguished by the interocular
space unarmed; and antennal segment I stout, and only basally constricted.
General distribution. Cited from Canada, Mexico, Panama and the United States (Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♀, Jarabacoa, 600 m, 11-
viii-1980, A. Norrbom (CMNH).
Merocoris (C.) typhaeus (Fabricius)

(Fig. 3)
Lygaeus typhaeus Fabricius, 1798: 537.
Discussion. Distinguished by having body thick, oblong-ovoid, thickly clothed with a mat of woolly pubescent
hairs; ocelli set on tubercles; interocular space with short tubercle near the inner border of each eye; antennal
segment I subclavate, tapering toward the base from the middle; anterolateral borders of pronotum with 3 to 4 large
teeth; humeral angles subacute; posterior margin deeply emarginate; and endocorium with large black glabrous
spot forming an irregular black patch.
General distribution. Cited from Cuba and the United States (Blöte 1938; Barber & Bruner 1947;
Baranowski & Slater 1986; Zayas 1988; Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♀, Santo Domingo,
Parque Mirador del Norte, 24-vi-2001, H. Takizawa (USNM).
Subfamily Coreinae
Tribe Acanthocerini
Acanthocerus Palisot de Beauvois 1818
Diagnosis. Body medium-sized, relatively robust, oblong. Antenniferous tubercles armed laterally with short and
stout spine. Antennal segment III cylindrical. Ocelli raised. Postocular tubercle exposed. Pronotum steeply
declivent. Anterolateral and posterolateral margins of pronotum uniformly dentate. Frontal angles projected into
stout spine, almost reaching the postocular tubercle. Humeri widely expended, slightly produced laterally with the
humeral spine blunt, directed outward and slightly backward. Posterior border straight. Triangular process present.
Dorsal surface of pronotum punctate, not tuberculate. Posterior lobe of metapleura bearing a lobe in the male.
Femora armed. Hind femora of males more incrasste than females and each dorsally tuberculate. Metathoracic
peritreme bilobate.
Distribution. The genus includes three species restricted to the Antilles: Cuba, the Dominican Republic, and
Haiti. The record of the genus in the United States (Florida) according to O’Shea (1975) and Baranowski & Slater
(1986) is probably incorrect due to mis-labeling.
Acanthocerus crucifer Palisot de Beauvois
(Fig. 4)
Acanthocerus crucifer Palisot, 1818, 204.
Discussion. Can be distinguished by having the body dorsally and ventrally dark reddish brown to dark castaneous
brown; anterolateral and posterolateral margins and posterior margin of pronotum, an incomplete longitudinal
stripe at midline of pronotal disk, claval vein, apical angles of corium and posterior margin of metapleura yellow;
dorsal abdominal segments shiny orange; and abdominal sterna IV–VI with an irregular yellow longitudinal stripe.
General distribution. Only known from Cuba and Hispaniola (Dominican Republic and Haiti).
HISPANIOLA: DOMINICAN REPUBLIC: Santo Domingo. HAITI: Port au Prince (Dallas 1852, Walker 1871,
Blöte 1938, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, Palacio de Engombe,
Santo Domingo Oeste, 18°27.047’N 69°59.802’W, 16-x-2012, D. Perez, S. Navarro (USNM); 1 ♀, Palacio de
Engombe, Santo Domingo Oeste, 18°27.047’N 69°59.802’W, 5-viii-2011, D. Perez, S. Navarro (USNM); 1 ♀,
HDR-021, Sendero Rabo de Gato, Parque Nacional Sierra de Bahoruco, Independencia Prov., 415 m, 18°19.042’N
71°34.279’W, 31-viii-2015, D. Perez, R. Bastardo, A. Sánchez. Day + UV light (USNM); 1 ♀, 2 km W San Rafael
del Yuma, nr. vertedero, 75 m, 18°26.276’N 68°41.520’W, D. Perez, R. Bastardo, RD-200 (USNM); 2 ♂♂, Prov.
La Altagracia, Higuey, Macao, 10-viii-1979, Mota, Reynoso (MNHNSD); 1 ♂, Prov. Salcedo, San José, 16-viii-
1979, Reynoso (MNHNSD); 1 ♂, Prov. La Vega, 25-ii-1974, Domínguez (MNHNSD); 1 ♀, Prov. La Altagracia, S.
R. Yuma, Guaraguao, 25-i-80, Marcano (MNHNSD); 4 ♂♂, 1 ♀, Monseñor Nouel, Bonao, Presa Rio Blanco, casa
de máquinas, 13-v-2001, C. Nuñez (MNHNSD); 1 ♂, 1 ♀, Alto del Rancho, Guaconejo, Prov. M. T. Sánchez, 170
m, 24-25-vii-2002, 19°18’75”N 69°56’66”W, D. Perez, B. Hierro, R. Bastardo (MNHNSD); 1 ♂, Prov. Santiago,
Tamboril, Guazumal Arriba, 4-xi-1979, Domínguez, Aquino (MNHNSD); 1 ♀, Prov. La Altagracia, Boca de
Yuma, Higuey, 5-vii-1979, Marcano (MNHNSD); 1 ♂, 1 ♀, Monseñor Nouel, 25.6 km ESE Maimón, 18°49’30”N
70°03’13”W, 110 m, 20-xi-2002, W. A. Zanol, C. W. Young, C. Staresinic, J. Rawlins (CMNH). Biological note:
Disturbed forest along stream. 3 ♂♂, La Altagracia, Higuey, Nisibón, 28-iv-1980, Reynoso y Mtz. (MNHNSD); 1
♂, Prov. Samaná, Samaná, Pto. Francés Viejo, 12-iv-1979, Domínguez (MNHNSD); 1 ♂, Salcedo, Los Azules,
Salcedo, 17-viii-79, Reynoso (MNHNSD); 1 ♀, Prov. Salcedo, Salcedo, San José, 16-viii-1979, Reynoso
(MNHNSD); 1 ♀, Barahona, eastern Sierra Bahoruco, Reserva Cachote, 12.8 km NE Paraiso, 18°05’54”N
71°11’21”W, 1230 m, 22-23-xi-2004, J. Rawlins, C. Young, V. Verdicic, C. Nuñez (CMNH). Biological note:
Cloud forest with tree ferns. 1 ♂, Elias Piña, north slope, Sierra de Neiba, 2 km SW Canada, 7 km WSW Hondo
Valle, 980 m, 18°42’N 71°45’W, 29-viii-1995, J. Rawlins, G. Onore, R. Davidson (CMNH); 7 ♂♂, 4 ♀♀,
Pedernales, along Rio Mulito, 13 km N Pedernales, 18°09’N 71°47’W, 230 m, 17-vii-1992, J. Rawlins, S.
Thomson, C. Young, R. Davidson (CMNH). Biological note: Riparian woodland. 2 ♂♂ 1 ♀, La Altagracia, Parque
del Este, caseta Guaraguao, 4.4 km SE Bayahibe, 18°19’59”N 68°48’42”W, 3 m, 26-27-v-2004, C. Young, J.
Rawlins, J. Fetzner, C. Nuñez (CMNH); 2 ♀♀, Pedernales, 1 km S Los Arroyos, 1125 m, 18°14’N 71°45’W, 18-x-
1991, R. Davidson, C. Young, S. Thompson, J. Rawlins (CMNH). Biological note: Second growth forest. 1 ♂, 2
♀♀, La Altagracia, 7 km NNE Boca de Chavón at Rio Chavón, 18°28’N 68°52’W, 20 m, 3-vii-1992, Rawlins,
Thompson, Young, Davidson (CMNH). Biological note: Riparian vegetation. 2 ♀♀, La Altagracia, 9.7 km NW
Punta Cana, 18°35’11”N 68°26’22”W, 36 m, 26.v.2004, Rawlins, Young, C. Nuñez (CMNH); 1 ♀, Pedernales, La
Abeja, 38 km NNW, Cabo Rojo, 18°09’N 71°38’W, 1250 m, 15-vii-1987, Rawlins, Davidson (CMNH); 1 ♀,
Santiago, Cordillera Septentrional, Pico Diego de Ocampo, summit, 19°35’06”N 70°44’23”W, 958 m, 31-v-2004,
Rawlins, Young, C. Nuñez, J. Fetzner (CMNH); 1 ♂, El Seibo, Loma Cocuyo, 6 km N Pedro Sánchez, 18°55’N
69°07’W, 475 m, 4-vii-1992, Young, Davidson, Rawlins, Thompson (CMNH). Biological note: Disturbed fields
and woodland. 1 ♂, San Lorenzo, 24-26-vi-1915 (UNAM). 1 ♂, 1 ♀, Peravia, Arroyo Cañas, 700 m, 5-8.viii.80, A.
Norrbom (UNAM); 2 ♂♂, 2 ♀♀, Jarabacoa, La Vega Prov., 1700’, vi-81, C. M. Stevens (UNAM); 3 ♂♂, La Vega,
Bamboo hole canyon, Rio Baiguate, 5 km SE Jarabacoa, 580 m, 22-vii-87, Davidson y Rawlins (UNAM); 1 ♂, La
Vega, NE Buena Vista, 24-31-v-1991, S. W. Dunkle (FSCA); 1 ♂, Pedernales Prov., 23 km N Cabo Rojo, 1200 m,
31-xii-86, Doyen y Santiago (UNAM); 1 ♂, 1 ♀, Sosua, E. Puerto Plata, 25.iii.84, M. Alfenito (UNAM); 1 ♂, 7-
15-i-84, M. Alfenito (UNAM); 1 ♂, Prov. Pedernales, Cabo Rojo, Alcoa Rd., 24-25-vii-85, G. Eickwort (UNAM);
1 ♂, Santo Domingo, iii.1907, H. H. Keays (UNAM); 2 ♂♂, 2 ♀♀, Azua, East side of crest Sierra Martín García, 7
km WNW Barrero, 860 m, 18°21’N 70°58’W, 25-26.viii.92, C. Young, R. Davidson, S. Thompson, J. Rawlins
(CMNH). Biological note: Cloud forest adjacent to disturbed forest. 1 ♂, 2 ♀♀, Pedernales, along Rio Mulito, 13
km N Pedernales, 18°09’N 71°46’W, 230 m, 17.viii.92, J. Rawlins, S. Thompson, C. Young, R. Davidson
(UNAM). Biological note: Riparian woodland. 1 ♂, 3 ♀♀, Rep. Dominicana, San Juan, 7 km N Arroyo Cano, 1
km S Los Frios, 1120 m, 18°52’N 71°01’W, 1-ix-95, J. Rawlins, G. Onore, R. Davidson (CMNH). Biological note:
Pine forest, second growth. 3 ♂♂, Pedernales, 19-20 km N Cabo Rojo, 375 m, 10-vii-96, R. Turnbow (UGAG); 1
♀, La Vega, 1 km W Manabao, 6-vi-94, R. Turnbow (UGAG). 1 ♀, Prov., El Seibo, 18.5 km W. El Seibo, 15.v.92,
R. Turnbow (UGAG); 1 ♀, Prov. Barahona, 4.5 km S Barahona, 22-v-92, R. Turnbow (UGAG); 1 ♂, 1 ♀, La Vega,
Constanza, 17-vii-86, R. Turnbow (UGAG); 2 ♂♂, 1 ♀, , Prov. Pedernales, 24 km N Cabo Rojo, 535 m, 11-vii-
1996, M. C. Thomas (FSCA); 1 ♀, Prov. La Altagracia, Nisibón, 6-v-78, Woodruff, Fairchild (FSCA); 8 ♂♂, 8
♀♀, La Altagracia, Higuey, Verón, 10-viii-79, Marcano, Mota, Reynoso (MNHNSD); 4 ♂♂, 4 ♀♀, 21-vii-79
(MNHNSD); 2 ♂♂, 1 ♀, Barahona Prov., 4.5 km S Barahona, 23-v-1992, M. C. Thomas (FSCA); 3 ♂♂, Prov.
Hato Mayor, Mango Limpio, 10-vi-1986, L. Stange, R. Woodruff (FSCA); 1 ♂, Hato Mayor, Juan Jiménez, 27-i-
1999, J. P. Michaud (UNAM). Biological note: On Citrus sinensis, sweet orange. 10 ♂♂, 10 ♀♀, Salcedo, Las
Cuevas, 29-viii-1979, Reynoso (MNHNSD); 1 ♂, 13.iv.1979 (MNHNSD); 1 ♀, Prov. Santiago, Tamboril, Licey al
Medio, 29-ix-1979, Marcano (MNHNSD); 1 ♀, Prov. Puerto Plata, La Cumbre, 28-xii-1978, Domínguez y Silfa
(MNHNSD); 1 ♀, Pedernales Prov., P. N. Jaragua, 3 km S of Los Tres Charcos, 17° 48’01”N 71°26’80”W, 99 m,
16-vi-2005, L. Chamorro, A. Konstantinov (UMSP); 1 ♂, La Vega Prov., P. N. Armando Bermúdez, 7 km SE
Jarabacoa, 970 m, 19°05’92”N 70°35’86”W, 23-v-2005, L. Chamorro, A. Konstantinov (UMSP); 1 ♂, La
Altagracia Prov., El Veron, rd. to Hoyo Azul, 18°33’72”N 68°26’87”W, 25-40 m, 1-vii-2005, L. Chamorro, A.
Konstantinov (UMSP); 1 ♀, La Altagracia Prov., P. N. del Este, Valle de Orqueta, 25 m, 18°22’93”N 68°46’63”W,
29-vi-2005, L. Chamorro, A. Konstantinov (UMSP); 1 ♀, Puerto Plata, 21-ii-1990, D. M. Webb (FMNH); 1 ♂, 2
♀♀, Barahona Prov., near Filipinas, Larimar Mine, 26-vi-6-vii-1992, F. W. Skillman jr. (CDFA); 4 ♀♀, Barahona
Prov., Baoruco mts., near Filipinas, Larimar Mine, 14-xii-1991, F. W. Skillman jr. (CDFA). Note: Beaten fresh
slash. 1 ♀, La Vega Prov., Jarabacoa, 9-viii-67, J. C. Schaffner (TAMU); 1 ♂, Peravia Prov., 21-viii-67, L. H.
Rolston (TAMU); 5 ♂♂, 6 ♀♀, Constanza, 2-6-vi-1969, Flint & Gomez (USNM); 1 ♂, 2 ♀♀, La Vaca, 6 km W. of
Jayaco, 2-vi-1969, Flint & Gomez (USNM); 1 ♂, 1 ♀, San Cristobal, vii-1969, J. Maldonado C. (USNM); 1 ♂, 1
♀, 8-vi-1995, Santo Domingo: Botanical Garden, 18.28 N 69.54 W, leg. Ehrman (USNM); 1 ♂, 2 ♀♀, La Vega
Prov., PN Armando Bermúdez, km 1-7 along trail W of La Ciénaga, 900-1100 m, 19° 01.753’N 70°54.654’W, 19-
vi-2010, S. Lingafelter (USNM); 1 ♂, 1 ♀, Paso de la Vaca, Mons. Nouel-Constanza Rd., Prov. La Vega, R. D., 27-
xii-1955. Alt. 1500 m, J. Maldonado Capriles, coll. (USNM); 1 ♂, 27-v-1995, Bani: at rd. nr 2, leg. Ehrman 18°17’
N 70°20’ W (USNM); 1 ♀, 27-v – 4 vi-1995, Barahona: Hotel Guarocuya, 18°13´ N 71°07’ W (USNM); 1 ♀, 10-
vi-1995, Santo Domingo Zool. Garden, 18.28 N 69.54 W leg. Ehrman (USNM); 1 ♂, La Vega: La Vega (3.5 km
NW Rio Camú, 19°13.8’N 70°35.2’W, 100 m, 18-v-1998, W. N. Mathis (USNM); 1 ♂, Wi, S. of Miches, 100-550
m, 15-ix-1973; Coll. #1, TJ Walker, JC Schuster (USNM); 5 ♂♂, 3 ♀♀, 4 mi. N. of Santo Domingo city, 22-vi-
1917 (359), Harold Morrison (USNM); 1 ♀, Duarte, Santo Domingo city, 21-vii-1917 (358), Harold Morrison
(USNM); 1 ♂, 6 ♀♀, 8 mi. up Macoris river, San Domingo, 16-vii- 1917, (354), Harold Morrison (USNM); 2 ♂♂,
2 ♀♀, At Blanton Mine, N. of San Christobal, 26-vii-1917, (376), Harold Morrison (USNM); 3 ♂♂, 4 ♀♀, Santo
Domingo, 6.8, WI, Aug. Busck Collector (USNM); 5 ♂♂, 2 ♀♀, Santo Domingo, 7-8, WI, Aug. Busck Collector
(USNM); 1 ♀, Santo Domingo, 3-iii-13, WI, P. G. Rossell Collector (USNM); 1 ♂, 2 ♀♀, San Francisco Mts.,
Santo Domingo, ix-15, WI, Aug. Busck Collector (USNM); 1 ♂, 1 ♀, Macoris, Santo Domingo, 3-iii-13, WI, P. G.
Russell Collector (USNM); 1 ♂, San Pedro de Macoris, 15-vii-1917, (335), Harold Morrison (USNM); 2 ♂♂, 1 ♀,
La Vega, Gualemati, S. D., vii-25, H. E. Box Coll. (USNM); 1 ♂, Santo Domingo, W.I., WLAbott Collector
(USNM); 2 ♀♀, Samana, Santo Domingo, Dr. Miller Coll. (USNM); 1 ♂, Romana, R. D., 4-xi-1913, W. V. Tover
Collector (USNM); 1 ♂, Jarabacoa, R. D., 14-vi-40, DeLeon Col. (USNM); 1 ♂, Barahona region, 1922, Collector
WLAbbott (USNM); 3 ♂♂, 4 ♀♀, La Altagracia Prov.: 2.7 km E. Batey Papagayo, elev. 150 ft., 18°55’24”N
68°44’21”W, 408 iv-2000, T. J. Henry & R. E. Woodruff (USNM). 2 ♂♂, La Vega Prov.: Rt. 12, 15 km SW of jct.
Rt. 1, above Bonao, el. 3900 ft., 19°00’36”N 70°33’35”W, 18-iv-2000, T. J. Henry & R. E. Woodruff. Taken from
Melastomataceae (USNM). 2 ♂♂, 1 ♀, Nisibón, El Cedro, 19-vii-1969, J. Maldonado C. (USNM); 1 ♂, La
Altagracia Prov., 9 xi 1962 (USNM); 2 ♂♂, 1 ♀, Santo Domingo Prov., Sierra Prieta, trail behind family house,
18°38.934’N 69°58.374’W, 140 m, M. Guidoti, D. Perez-Gelabert (USNM); 1 ♂, Monseñor Nouel Prov., rd. to
Blanco hydroelectric, 18°52.940’N 70°30.346’W, 700 m, 13-viii-2016, D. Perez, R. Bastardo, A. Sanchez
(USNM);1 ♂, Monseñor Nouel Prov., Bonao, old field with gate inside town, 18°55.624’N 70°25.701’W, 250 m,
M. Guidoti, D. Perez-Gelabert (USNM); 2 ♂♂, Santiago Prov., Trail to Monte Diego de Ocampo, 750-800 m, 10-
vii-2015, 19°34.326’N 70°43.786’W, D. Perez, S. Navarro (day + uv light) (USNM); 1 ♂, 1 ♀, San Pedro de
Macorís Prov., Juan Dolio, forest on north side of rd., 18°25.165’N 69°28.059’W, nr sea level, 3-viii-2016, D.
Perez, S. Navarro, J. Borbón (USNM); 1 ♂, Santiago Prov., Babosico, 500 m, 19°20.983’N 70°37.447’W, 12-vii-
2015, D. Perez, S. Navarro (day) (USNM); 1 ♀, San Cristóbal Prov., La Colonia to Los Cacaos, 2 km SE Los
Cacaos, 550 m, 18°31.444’N 70°16.835’W, 24-vii-2017, T. Henry, M. Guidoti, D. Perez-Gelabert (USNM); 1 ♀,
DR-2016-005 San Cristóbal Prov., by Nizao river (first stop), nr. hydroelectric, 18°26.044’N 70°15.656’W, 530 m,
10-viii-2016, D. Perez, C. de Soto Molinari, D. Pou (USNM). HAITI: 1 ♂, Hinche, Haiti, 26-VIII-1924, G. N.
Wolcott Coll. (USNM) ; 2 ♀♀, Port au Prince, 24-viii-1924, G. N. Wolcott Coll. (USNM); 1 ♀, Pt. Prince, iv-25,
Collection O. Hoffman (USNM); 1 ♂, Haiti, J. B. Terris Coll. (USNM); 1 ♂, Haiti, No. -2, Fond-des-Negres, 12-
vi-1930, H. L. Dozier Collector (USNM); 1 ♂, Hinche, 18-v-1925, W. A. Hoffman (USNM); 1 ♂, Haiti (UNAM).
1 ♀, Chatard, Dept. du Nord, 21-v-1959, M. W. Sanderson (FMNH) ; 1 ♂, 2 ♀♀, Furcy, 15-v-1959, M. W.
Sanderson, T. H. Farr (CASC) ; 2 ♂♂, 2 ♀♀, Port-au-Prince, SW edge 7-ix-73, Coll. #1, T. J. Walker, J. C.
Schuster (USNM).
Tribe Anisoscelini
Key to Hispaniola genera and species of Anisoscelini
1. Hind tibiae foliaceous, strongly expanded on both sides . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

- Hind tibiae simple, not expanded . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
2. Antennal segment I much longer than total length of head; antennal segments II–III with or without flat lateral expansions . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Chondrocera Laporte … Ch. mirabilis Brailovsky & Barrera
- Antennal segment I equal to total length of head or slightly longer; antennal segments II–III simple, not expanded . . . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Leptoglossus Guerin
3. Body surface scattered with metallic iridescence . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
- Body surface entirely rather dull . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
4. Posttylar depression absent; hind femora dorsally tuberculate; humeral angles remarkably expanded into large and acute spine
directed outward and forward; calli with two conspicuous projections (less developed in females) each lateral to midline. . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Rhytidophthia Brailovsky … R. splendida Valdes
- Posttylar depression forming single sulcus; hind femora dorsally smooth; humeral angles tapering into short medium size acute
to subacute spine, directed outward and never forward; calli without spine-like projections . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Phthia Stål … P. rubropicta (Westwood)
5. Rostrum longer, reaching posterior border of abdominal sternite III or anterior border of IV; antennal segment I slender, longer
than 2.70 mm; antennal segment IV longer than 4.20 mm; scutellum, clavus, corium and hind femora without granules; tylus
not raised above jugum . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Phthiacnemia Brailovsky … P. picta (Drury)
- Rostrum reaching middle third of metasternum; antennal segment I stout, shorter than 2.20 mm; antennal segment IV shorter
than 2.50 mm; scutellum, clavus, corium and hind femora densely granulate with yellowish white granules; tylus compressed
above jugum but never projecting upward in the form of a triangular spine . .Mamurius Stål* … M. cubanus Barber & Bruner
*This genus currently considered as incertae sedis does not belong to Nematopodini or Anisoscelini and has been considered
elsewhere (O’Shea 1979, 1980). As long as it precise tribal position is not ascertained, and in order to facilitate its
identification, in this contribution it is temporarly assigned to that tribe.
Chondrocera Laporte 1832
Diagnosis. Hind tibiae with inner and outer face expanded. Head non declivent, porrect, extending forward beyond
antenniferous tubercles. Antennal segments II–III expanded and flattened or nearly cylindrical. Posttylar
depression single. Humeral angles projected on a short to medium-sized spine directed laterally and slightly
backward.
Distribution. The genus includes two species and one subspecies cited from the Bahamas, Cuba, the
Dominican Republic, Guadeloupe and the United States. One species occurs in Hispaniola (Baranowski & Slater
1986, Brailovsky 1994, Packauskas 2010).
Chondrocera mirabilis Brailovsky & Barrera
(Fig. 5)
Chondrocera mirabilis Brailovsky & Barrera, 1994: 54–57.
Discussion. Can be distinguished from the other species in the genus by having the pronotum, scutellum, clavus
and corium with bluish green metallic iridescences; antennal segments II–III cylindrical; calli transverselly raised;
and apex of scutellum yellow.
General distribution. Known from the Dominican Republic and Guadaloupe. HISPANIOLA. DOMINICAN
REPUBLIC: San Pedro Macoris, y La Vega (Brailovsky & Barrera 1994, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, 1 ♀, HDR-001,
Parque Mirador del Norte, Santo Domingo, 18°517393’N 69°17912’W, 9-vii-2015, D. Perez, S. Navarro (USNM);
1 ♂, HDR-014, La Charca, km 28 rd. to El Aceitillar, Pedernales Prov., 798 m, 18°06.750’N 71°37.088’W, 25-vii-
2015, D. Perez, J. Espinal (Day + UV light) (USNM); 1 ♂, Boca de Yuma, P. N. del Este, La Altagracia Prov., 20
m, 18°21.875’N, 68°37.081’W, 16–17-xii-2003, D. Perez, R. Bastardo, RD-199 (USNM); 1 ♂, 1 ♀, Rd. to
Monteada Nueva, N. of Polo, Barahona Prov., 18°06.985’N 71°15.246’W, 810 m, 28-viii-2014, D. Perez, C. de
Soto M. (USNM); 1 ♀, Entrance to Playa Cumayasa, San Pedro de Macorís Prov., 18-xii-2003, D. Perez, B. Hierro,
R. Bastardo, RD-206 (USNM); 1 ♀, Pedernales Prov., 25.5 km N Cabo Rojo, 20-v-1992, R. Turnbow (UGAG); 1
♀, Prov. Barahona, Filipinas, Larimar Mine, 13-xii-91, R. E. Woodruff (FSCA); 1 ♂, Distrito Nacional, Boca
Chica, 20-ix-1979, Mota, Domínguez (MNHNSD); 1 ♀, San Cristóbal Prov., Haina, 13-vii-1980, Domínguez
(MNHNSD); 1 ♂, Barahona, 4.5 km Sur Barahona, 4 km W Hwy 2, 17-v-1992, M. C. Thomas (UNAM); 1 ♂, 1 ♀,
Barahona, 9.2 km NW Paraiso, confluence del Río Nizao y Río Coltico, 18-03N 71-12W, 230 m, 9–10-viii-90,
Rawlins, Thompson (UNAM); 2 ♀♀, Barahona, 5 km SE Polo, slopes of Loma La Torre, 18-03N 71-16W, 980 m,
18-vii-92, Young, Davidson, Thompson, Rawlins (UNAM); 1 ♀, Puerto Plata Prov., Sosua, 19-vii-86, M. Carlon
(UNAM); 1 ♂, Prov., S. F. Macorís, 27-xii-67, L. H. Rolston (UNAM); 1 ♀, Prov. La Vega, 8-viii-67, L. H. Rolston
(UNAM); 1 ♀, San Pedro de Macorís, 18-v-83, E. Giesbert (UNAM); 1 ♀, Hato Mayor, Parque Los Haitises, E. of
Trepada Alta, 12 km W El Valle, 18-59N 69-30W, 145 m, 6-vii-92, J. Rawlins, S. Thompson, C. Young, R.
Davidson (UNAM). Biological note: Mesic forest on limestone. 1 ♀, Azua, east side of crest Sierra Martín García,
7 km WNW Barrero, 18°21’N 70°58’W, 860 m, 25–26-vii-92, C. Young, R. Davidson, S. Thompson, J. Rawlins
(UNAM). Biological note: Cloud forest adjacent to disturbed forest. 1 ♀, La Altagracia, 2 km N Bayahibe,
18°23’N 68°51’W, 10 m, 3-vii-92, C. Young, R. Davidson, S. Thompson, J. Rawlins (UNAM). Biological note:
Dry seasonal forest on limestone. 1 ♂, San Pedro Prov., 13 km E Boca Chica, 14-v-1992, R. Turnbow (UGAG); 1
♀, Bosque húmedo camino a Angel Felix, 19-viii-2014, 18.626900 -71.722925, M. R. Bobadilla (IIBZ).
Leptoglossus Guérin-Méneville 1831
Diagnosis. Medium to large, elongate species. Head porrect, longer than wide, shorter than length of pronotum and
extending forward beyond antenniferous tubercles. Tylus slightly exceeding juga. Posttylar depression double.
Humeral angles obtuse or expanded into acute projections. Femora ventrally armed with two rows of distal teeth.
Hind femora swollen, usually thicker in males, and always dorsally spined or tuberculated. Hind tibiae dilated,
frequently strongly and leaf-like with outer dilation wider than inner one.
Distribution. A large genus with 61 species and one subspecies widely distributed from southern Canada,
throughout the United States, Mexico, Central America, Greater and Lesser Antilles, and South America (Allen
1969, Baranowski & Slater 1986, Packauskas 2010, Brailovsky 2014). Four species occur in Hispaniola.
Key to Hispaniola species of Leptoglossus
1. Thoracic pleura with 10 to 12 strongly contrasting yellowish maculae; thoracic and abdominal sterna without numerous black
discoidal spots; pronotal disk with narrow, arcuate, pale yellowish transverse fascia; hind tibiae with outer dilation extending
for 85–90% the maximal length of tibiae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .L. gonagra (Fabricius)
- Thoracic pleura orange to pale yellowish or dark reddish brown, without strongly contrasting yellowish markings; thorax and
abdominal sterna with or without numerous small black dots; pronotal disk lack a narrow yellowish transverse fascia; hind tib-
iae with outer dilation extending for at most 70% of the maximal length of tibiae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
2. Yellow transverse fascia on corium always present, rectangular, with anterior and posterior margins straight and parallel . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . L. balteatus (Linnaeus)
- Yellow transverse fascia on corium irregular (zig-zag) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
3. Pronotal disk unicolorous, black to dark brown, without contrasting yellowish areas; middle third of male genital capsule
straight . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . L. concolor (Walker)
- Pronotal disk with a yellowish rectangular to quadrangular mark not reaching the anterolateral margins; middle third of male
genital capsule widelly open . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . L. confusus Alayo & Grillo
Leptoglossus balteatus (Linnaeus)

(Fig. 6)
Cimex balteatus Linnaeus, 1771: 534.
Discussion. This species is known by having the thoracic pleura pale yellowish, without contrasting yellowish
markings; thoracic and abdominal sterna with with numerous small black dots; a yellowish transverse fascia on
corium rectangular, with anterior and posterior margins straight and parallel; posterior margin of pronotum yellow;
pronotal disk with a pair of large yellow spots sometimes extensive and coalescing mesally; antennal segments II
and III orange with apical third reddish; and humeral angles acuminate, acute and obliquely ascending.
General distribution. Known only from the Greater Antilles including the Bahamas, Cuba, the Dominican
Republic, Grenada, Haiti, Jamaica, Puerto Rico, St. Thomas the United States (Florida). HISPANIOLA:
DOMINICAN REPUBLIC: San José las Matas, and Puerto Plata. HAITI: Any formal records are included (Barber
1939, Allen 1969, Baranowski & Slater 1986, Brailovsky 2014, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, 1 ♀, La Altagracia
Prov., Punta Cana, 26-xii-2008, D. Perez-Gelabert (USNM); 1 ♂, 1 ♀, Road down from door, 800–1000 m, Parque
Nacional Sierra de Bahoruco, Independencia Prov., 12.xii.2003, D. Perez, R. Bastardo, B. Hierro, RD192 (USNM);
1 ♂, Rio Higuamo, San Pedro de Macorís Prov., 5-ix-2004, C. de Soto M. (USNM); 1 ♂, Pedernales Prov., Cabo
Rojo, Alcoa Rd., km 33, 27-vii-85, G. Eickwort (UNAM); 1 ♀, Monte Cristi, 5 km N Villa Elisa, 3-vi-1994, R.
Turnbow (UGAG); 1 ♀, Barahona prov., 4. 5 km S of Barahona, 17-v-1992, R. Turnbow (UGAG); 1 ♂, Prov.
Pedernales, Cabo Rojo, 21-v-1992, R. Turnbow (UNAM). Biological note: at light. 1 ♂, La Altagracia, 2 km N
Bayahibe, 18°23’N 68°31’W, 10 m, 3-vii-92, C. Young, R. Davidson, S. Thompson, J. Rawlins (CMNH).
Biological note: Dry seasonal forest on limestone. 1 ♀, Haina-San Cristóbal, 16-xii-1982, H. Domínguez
(UNAM); 1 ♂, Prov. Baní, Galeón, Peravia, 22-viii-1979, Domínguez – Mota (MNHNSD); 1 ♀, Pedernales, Cabo
Rojo, 10 m, 17°55’N 71°39’W, 26–27-ix-1991, Young, Thompson, Davidson, Rawlins (CMNH); 1 ♂, Isla Beata,
21–22-x-1978, Marcano (MNHNSD); 2 ♂♂, 1 ♀, La Altagracia Prov., Parque del Este, 2.9 km SW Boca de Yuma,
18°21’51”N 68°37’05”W, 11 m, 28-v-2004, J. Rawlins, C. Young, C. Nuñez, J. Fetzner (CMNH). Biological note:
Semihumid dry forest. Limestone, UV light, also hand collecting. 1 ♂, 2 ♀♀, La Altagracia Prov., Parque del Este,
caseta Guaraguao, 4.4 km SE Bayahibe, 18°19’59”N 68°48’42”W, 3 m, 26–27-v-2004, Rawlins, Young, Fetzner,
Nuñez (CMNH); 1 ♂, Prov. Montecristi, RD-089, 8 km E Montecristi, 19°48’93”N 71°35’63”W, 10-xii-2002, D.
Perez, R. Bastardo (MNHNSD); 2 ♂♂, San Juan Prov., San Juan, 16-viii-1973, Domínguez (MNHNSD); 1 ♂, 1 ♀,
San Cristóbal Prov., San Cristóbal, 17-viii-1980, Marcano (MNHNSD); 1 ♂, 20-vii-80 (MNHNSD). 1 ♂, Santo
Domingo (UNAM); 1 ♂, Pedernales Prov., Cabo Rojo, Alcoa Rd., km 33, 27-vii-1985, G. Eickwort (UNAM); 1 ♂,
Pedernales, Prov., Cabo Rojo, 21-v-1992, R. Turnbow (UNAM); 1 ♀, San Pedro de Macoris Prov., Juan Dolio,
forest on north side of rd., 18°25.165’N 69°28.059’W, nr sea level, 2-viii-2016, D. Perez, S. Navarro, J. Borbón
(USNM); 1 ♂, 27-v-1995, DomR, BaniL at rd. nr. 2, leg. Ehrman, 18.17° N, 70.20° W (USNM); 1 ♀, 27-v -4-vi-
1995, Barahona: Hotel Guarocuya, 18.13° N 71.07° W (USNM); 2 ♀♀, Macoris, Santo Domingo, 3-iii-13, WI, P.
G. Russell Collector (USNM); 1 ♀, Peravia, Matadero, (18°24’8.2”N 70°25’12.2”W), 480 m, 1-viii-2005, Luz
negra, coll. S. Vélez, J. Henríquez, A. Marmolejo, R. Bastardo (IIBZ). HAITI. 1 ♀, Puerto Principe, (UNAM); 1 ♂,
Port au Prince, 27.viii.1901, G. Kritel (ZIZM); 1 ♀, Hinche, 19-xii-1930, H. L. Dozier (USNM); 1 ♀, Hinche, 19-
xii-1930, H. L. Dozier Collector (USNM); 1 ♂, Port-au-Prince, ii-1925 G. N. Wolcott (USNM).
Leptoglossus concolor (Walker)
(Fig. 8)
Anisoscelis concolor Walker, 1871: 128.
Discussion. The summary of characters separate this species from the other known Leptoglossus of Hispaniola:
thoracic pleura pale yellow to yellowish orange, without contrasting yellowish markings; thoracic and abdominal
sterna with numerous small black dots; yellowish transverse fascia on corium irregular (zig-zag); pronotum with
anterolateral margins entire; posterolateral margins of pronotum serrate; pronotal disk dark brown to dark reddish
brown without contrasting yellowish marks; hind tibial dilations conspicuously phylliform; hind tibiae with outer
dilations occupying less than 80% of the maximal length of the tibiae; antennal segment I bicolorous; rostrum
extending well onto abdomen; corial veins unicolorous with the surface; and pronotal calli with rough surface
texture, numerous dark thick hairs interspersed with pale hairs.
General distribution. Distributed widely in Mexico, Central America, Greater Antilles and south of the
United States. British Honduras, Costa Rica, Cuba, the Dominican Republic, Guatemala, Haiti, Panama, Puerto
Rico, the United States and Virgin Islands. HISPANIOLA: DOMINICAN REPUBLIC: Puerto Plata and Sánchez.
HAITI: St. Marc. (Allen 1969, Baranowski & Slater 1986, Packauskas 2010, Brailovsky 2014).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♂♂, 1 ♀, Santiago,
Rincón de Piedra, S. J. de las Matas, 6-i-1979, Marcano (MNHNSD); 2 ♂♂, 1 ♀, La Vega 1 km W Manabao, 6-vi-
1994, R. Turnbow (UGAG); 1 ♂, 1 ♀, La Vega near mouth Arroyo Los Dajaos, 5 km E Manabao, 740 m, 19°04’N
70°45’W, 9-x-1991, J. Rawlins, R. Davidson, C. Young, S. Thompson (CMNH). Biological note: Riparian
woodland. 4 ♂♂, 4 ♀♀, Haina, San Cristóbal, 16-xii-1982, H. Domínguez (UNAM); 1 ♂, 1 ♀, Prov. Santiago,
Licey al Medio, Tamboril, 29-ix-1979, Marcano (MNHNSD); 1 ♀, Prov. San Cristóbal, Haina, S. Cristóbal, 20-vii-
1980, Domínguez (MNHNSD); 1 ♀, La Vega Prov., 4 km E. of La Ciénaga de Manabao, 3050 ft., 19°04’47”N
70°49’29”W, 19-iv-2000, T.J. Henry & R. E. Woodruff, black lights (USNM).
Leptoglossus confusus Alayo & Grillo

(Fig. 7)
Leptoglossus confusus Alayo & Grillo, 1977: 97.
Discussion. The following characters differentiate this species from the remaining members of the genus
Leptoglossus present in Hispaniola: thoracic pleura pale yellow to yellowish orange, without contrasting yellowish
markings; thoracic and abdominal sterna with numerous small black dots; yellowish transverse fascia on corium
irregular (zig-zag); pronotum with anterolateral margins entire; posterolateral margins of pronotum serrate; hind
tibial dilations conspicuously phylliform; posterior lobe of pronotal disk with contrasting yellowish rectangular
transverse band not reaching lateral margins; hind tibiae with outer dilations occupying less than 80% of the total
length of tibiae; and antennal segment I bicolorous.
General distribution. Formerly only known from Cuba (Alayo & Grillo 1977, Packauskas 2010, Brailovsky
2014). This species is cited for the first time from Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, Haina—San Cristóbal,
16-xii-1982, H. Domínguez (UNAM); 2 ♀♀, Santiago Prov., Licey al Medio, Santiago, 8-xii-1979, L. Marcano
(MNHNSD); 2 ♂♂, 1 ♀, Prov. San Cristóbal, Campus Loyola, 5-iii-1976, E. Marcano, #19644 (IIBZ); 1 ♂, 2 ♀♀,
Prov. San José de Ocoa, Bejucal, Los Polos Mojados, 11-iv-1976, E. Marcano & J. Cicero, #19725 (IIBZ); 1 ♂, 1
♀, Prov. San Cristóbal, Campus Loyola, 5-iii-1976, E. Marcano, #19645 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus
Loyola, 5-iii-1976, E. Marcano, #19655 (IIBZ); 2 ♂♂, 4 ♀♀, Prov. San Cristóbal, Campus Loyola, 10-iii-1976, E.
Marcano, #19688 (IIBZ); 1 ♂, Prov. San José de Ocoa, Bejucal, Los Polos Mojados, 11-iv-1976, E. Marcano & J.
Cicero, #19724 (IIBZ); 1 ♂, Prov. Santiago, Tamboril, Licey al Medio, 28-xii-1976 E. Marcano, #20966 (IIBZ); 1
♀, Prov. Monte Cristi, xii-1969, E. Marcano, #5495. sobre Guayaba (IIBZ); 1 ♂, Prov. San Cristóbal, Campus
Loyola, 10-iii-1976, E. Marcano, #19695 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 6-vi-1971, E.
Marcano, #10298 (IIBZ); 1 ♂, Prov. Santo Domingo, Los Minas, 24-x-1965, E. Marcano, #2610 (IIBZ); 1 ♂, Prov.
La Vega, Jarabacoa, Manabao, Los Tablones, 28-iii-1964, E. Marcano, #442; 2 ♀♀, Prov. San Cristóbal, Campus
Loyola, 5-iii-1976, E. Marcano, #19651 (IIBZ); (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 10-iii-1976, E.
Marcano, #19696 (IIBZ); 1 ♀, Prov. San Cristóbal, 19-ii-1975 E. Marcano #17610 (IIBZ); 1 ♀, Santo Domingo,
Los Minas, 27-Iix-1973 E. Marcano #14155 (IIBZ); 1 ♀, Prov. Monte Plata, Bayaguana, Hidalgo, Pilancón, 19-x-
1974 E. Marcano #17272 (IIBZ); 1 ♀, Prov. Monte Cristi, xii-1969, E. Marcano, #5498. sobre Guayaba (IIBZ); 1
♀, Prov. Santiago, Tamboril, Licey al Medio, 28-xii-1976 E. Marcano, #20961 (IIBZ); 1 ♀, Engombe, Santo
Domingo, D. N. 6-iii-76, UASD, Guayaba (IIBZ); 1 ♀, Prov. Santo Domingo, Engombe, 19-ix-2005, R. Rodríguez
(IIBZ); 1 ♀, Prov. Santo Domingo, Parque Mirador del Sur, 11-v-2017, S. García (IIBZ); 1 ♀, Prov. Sánchez
Ramírez, Cotuí, Cevicos, Los Tamarindos, 14-ii-2015, D. Sosa (IIBZ); 1 ♂, Prov. San Cristóbal, Cambita,
Garabito, 14-ii-2017, A. Ramos (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 5-iii-1976, E. Marcano, #19667
(IIBZ); 2 ♂♂, Prov. San Cristóbal, Campus Loyola, 10-iii-1976, E. Marcano, #19689 (IIBZ).
Leptoglossus gonagra (Fabricius)

(Fig. 9)
Cimex gonagra Fabricius, 1775: 708.
Discussion. A unique set of chromatic characters which identifies this species are the thoracic pleura with at least
10 strongly contrasting yellowish maculae, abdominal sterna with six to seven complete or maculate yellowish
longitudinal fascia and pronotal disk with narrow arcute, pale yellowish transverse fascia. Additionaly the hind
tibiae with outer dilation extending for 85–90% the maximal length of tibiae.
General distribution. A wide-ranging species occurring throughout the south of the United States, Mexico,
Central America, South America, Greater Antilles, and introduced in Africa, Australia, Canary Islands, Ceylon,
Cook Islands, Fiji, India, Indonesia, Madagascar, Malay archipelago, Micronesia, New Hebrides, New Caledonia,
New Guinea, Phillipines, Samoa, Seychelles, Solomon Islands, South East Asia, Tahiti, and Taiwan. In the Western
Hemisphere this species is cited from Argentina, Bolivia, Brazil, British Honduras, Cuba, the Dominican Republic,
El Salvador, Haiti, Jamaica, Mexico, Nevis, Nicaragua, Paraguay, Puerto Rico, St. Croix, St. Vincent, the United
States, Uruguay, Venezuela and Virgin Islands. HISPANIOLA: DOMINICAN REPUBLIC: Puerto Plata and
Sánchez. HAITI: Grande Rivière and Cape Haytien (Barber 1939, Allen 1969, Baranowski & Slater 1986,
Packauskas 2010, Brailovsky 2014).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♂♂, San Cristóbal Prov., 3
km N La Colonia, Mano Matuey, 7-ix-2008, D. Perez, B. Hierro, R. Bastardo, S. Medrano (USNM); 1 ♂, Palacio
de Engombe, Santo Domingo Oeste, 18°27.047’N 69°59.802’W, 5-viii-2011, D. Perez, S. Navarro (USNM); 1 ♀,
San Cristóbal Prov., 3 km N La Colonia, Mano Matuey, 7-ix-2008, D. Perez, B. Hierro, R. Bastardo, S. Medrano
(USNM); 1 ♂, Prov. Azua, El Número, Azua, 22-vii-1979, Domínguez, Marcano, Reynoso (MNHNSD); 1 ♀,
Prov. Independencia, La Azufrada, Descubierta, 6-vii-1979, Marcano (MNHNSD); 1 ♂, Prov. San Cristóbal,
Haina, 13-vii-1980, Domínguez (MNHNSD); 1 ♀, Barahona Prov., Filipinas, Larimar mine, 26-vi–6-vii-1992, F.
W. Skillman, jr. (UNAM); 1 ♀, Prov. La Vega, 10-iv-1977, Domínguez (MNHNSD); 2 ♂♂, Prov. San Cristóbal,
San Cristóbal, Haina, 28-vi-80, Domínguez (MNHNSD); 1 ♂, 1 ♀, Prov. Peravia, Baní, El Recodo, 10-xii-1978,
Domínguez (MNHNSD); 1 ♀, Finca Goya, 18-vii-90, R. Barnum (UNAM); 1 ♀, La Altagracia, 2 km S.W. campo
Nuevo, 1-viii-1979, L. B. O’Brien (UNAM); 6 ♂♂, 2 ♀♀, Prov. Azua, El Número, 22-vii-1979, Domínguez,
Marcano, Reynoso (MNHNSD); 1 ♂, Prov. Peravia, Bani, La Angostura, 15-ix-1979, Domínguez, Reynoso
(MNHNSD); 1 ♀, Prov. Azua, Tábara Abajo, 22-xii-1979, Marcano, Mtz. (MNHNSD); 1 ♀, Distrito Nal.,
Engombe, 20-ix-1979, Reynoso (MNHNSD); 1 ♀, San Cristóbal, Los Desamparados, 20-ix-88, Gómez (UNAM);
1 ♂, , Prov. Puerto Plata, La Cumbre, 6-viii-1974 E. Marcano #16753 (IIBZ); 2 ♂♂, 1 ♀, Prov. San Cristóbal,
Campus Loyola, 10-iii-1976, E. Marcano, #19691 (IIBZ); 1 ♂, 3 ♀♀, Prov. San Cristóbal, Campus Loyola, 10-iii-
1976, E. Marcano, #19692 (IIBZ); 1 ♂, Prov. San Cristóbal, Cambita, 8-vi-1976, E. Marcano, #20092 (IIBZ); 1 ♂,
Santo Domingo Este, El Almirante, 15-v-2017, V. Lépez (IIBZ); 1 ♀, Prov. María Trinidad Sánchez, Cabrera,
Arroyo Salado, Baoba del Pinar, 2-xi-1975, E. Marcano & J. Cicero, #19182 (IIBZ); 1 ♀, Prov. San José de Ocoa,
La Horma, 15-v-1976, E. Marcano & A. Abud, #20182 (IIBZ); 1 ♀, Prov. Santo Domingo, 8-ii-1970, E. Marcano,
#6007 (IIBZ); 1 ♀, Prov. San Cristóbal, Cambita, 8-vi-1976, E. Marcano, #20093 (IIBZ); 1 ♀, Prov. La Romana,
Batey Diego, 18-vi-1970, E. Marcano & R. Hansen, #7653 (IIBZ); 1 ♀, Santo Domingo, Los Minas, 27-ix-1973,
E. Marcano #14153 (IIBZ); 1 ♀, Santo Domingo Norte, Guanuma, 20-iii-2015, Juan C. Nuñez (IIBZ); 1 ♀, Santo
Domingo Norte, Guanuma, 24-x-2014, Ana Maria (IIBZ); 1 ♀, Engombe, Santo Domingo, D. N., 4-iii-76, UASD,
Auyama (IIBZ). HAITI: 1 ♂, Port Prince, iv-25, collection O. Hofmann (USNM).
Mamurius Stål 1862
Diagnosis. Body less than 15 mm, robust, stout and non-flattened dorsoventrally. Antennal segment I tuberculate.
Antennal segments II–III cylindrical not dilated. Antennal segment IV longer or subequal to III. Antenniferous
tubercles small, widely separated. Tylus tuberculate, projecting anteriorly to antenniferous tubercles. Pronotum
steeply declivent. Callar region flat. Frontal angles with large nodulose projection. Humeral angles slightly raised
above posterior lobe of pronotal disk, produced laterally, and ending in a short stout spine. Anterolateral margins
armed with large teeth. Mesosternum lacking median furrow. Femora ventrally armed. Hind femora incrassate in
both sexes, more in males, armed distally with large spines. Fore and middle tibiae sulcate, unarmed. Hind tibiae
flattened, inner margin armed with a row of small teeth in both sexes.
Distribution. Small genus with two known species cited from Cuba, Curacao, Mexico, Nicaragua, and the
United States (Barber & Bruner 1947, O’Shea 1979, Brailovsky 1982, Packauskas 2010). The genus is cited for the
first time from Hispaniola.
Mamurius cubanus Barber & Bruner

(Fig. 10)
Mamurius cubanus Barber & Bruner, 1947: 78–79.
Discussion. Body size less than 10 mm. Total length of male antennal segments: I, 1.42, II, 1.73, III, 1.48, IV, 1.56
(female unknown) (versus I, 1.92–2.23, II, 2.54–2.91, III, 2.04–2.35, IV, 2.17–2.48 on Mamurius mopsus Stål. First
measurement male, second female). Costal border of corium tuberculate. Connexival segments III–VI black to dark
brown with anterior border yellow and middle third of upper border with small yellow spot. Connexival segment
VII dark castaneous orange with upper border black with two yellow discoidal spots. Dorsal abdominal segments
shiny orange.
General distribution. Known from Cuba and Mexico (Barber & Bruner 1947, O’Shea 1979, Brailovsky 1982,
Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1♂, Barahona Prov.,
Masaco, S of Polo, 812 m, 18°02.495’N–71°18.408’W, (UV Light), 28-viii-2014, D. Perez, C. de Soto M.
(USNM). 1♂, Barahona Prov., rd. Barahona-Paraiso, off old field, 75 m, 18°11.435 N– 71°05.072W, 29.vii.2017,
T. Henry, M. Guidoti, D. Perez (USNM). 1♂, Peravia Prov., rd-198 Matadero, start of hills, 450 m, 18°24.464´N-
70°25.736W, 14-xii-2003, D. Perez, B. Hierro, R. Bastardo (USNM). 1♂, San Juan, 15 km E San Juan, 8-viii-1984,
C. W. O’Brien (UNAM). 1♂, S. R. 14 km E of Santo Rodriguez, 20-v-1978, C. W., & L. B. O’Brien and Marshall
(UNAM). 1 ♀, Peravia Prov., S. of El Matadero, NW of Baní, 400 m, 25-vii-2017, 18°23.489’N 70°25.855’W, T.
Henry, M. Guidoti, & D. Perez, ex. Melochia tomentosa (USNM). 1 ♂, Pedernales Prov., near Oviedo,
17°50.503’N, 071°24.425’W, 160 m, 1-viii-2017, T. Henry, M. Guidoti, D. Perez (USNM); 1 ♂, Prov. Montecristi,
Hatillo Palma, 29-xi-1964, E. Marcano, #912 (IIBZ); 1 ♀, Prov. Santiago, Jánico, La Jagua, 11-x-1975, E. Marcano
& J. Cicero, #19019 (IIBZ).
Phthia Stål 1862
Diagnosis. Medium size, elongate. Body surface scattered with metallic iridescence. Dorsal surface of head
entirely rather dull. Posttylar depression forming a single sulcus. Calli flat. Anterolateral borders of pronotum
smooth. Posterior lobe of pronotal disk with low longitudinal carinae. Humeral angles tapering into short or
medium size subacute spine, directed outward and never forward. Metathoracic peritreme bilobate. Hind femora
dorsally smooth.
Distribution. A small genus with six species, represented throughout the Neotropics (Brailovsky 2009,
Packauskas 2010). The genus is cited for the first time from Hispaniola.
Phthia rubropicta (Westwood)

(Fig. 11)
Leptoscelis rubropicta Westwood, 1842: 17.
Discussion. An attractive species characterized by having the antennal segment I longer than head; head and
pronotum anteriorly shiny bluish; anterior lobe of pronotal disk with narrow, arcuate, yellowish transverse fascia;
thoracic pleura with at least three strongly contrasting rectangular to quadrate maculae; mesopleura and metapleura
without black medial macula; hind tibiae simple; abdominal sterna III–VI lateral to midline with yellowish
maculae; and metallic iridescence restricted to ventral surface of head, calli, thorax, coxae, dorsal abdominal
segments III–VI and abdominal sterna III–VII.
General distribution. Cuba, Guadeloupe, Puerto Rico, and St. Vincent (Westwood 1842, Barber 1939, Barber
& Bruner 1947, Brailovsky 2009, Packauskas 2010). This species is cited for the first time from Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 5 ♂♂, 6 ♀♀, Santo
Domingo, D.N., 13 km Este, 2-ii-83, H. Domínguez (UNAM). Biological note: on Pisidium guajaba, and,
Cucurbita pepo (Cucurbitaceae) ; 1 ♂, Prov. Santo Domingo, 8-ii-1970, E. Marcano, #6015 (IIBZ); 1 ♂, Prov.
Santo Domingo, 8-ii-1970, E. Marcano, #6016 (IIBZ); 1 ♂, Prov. Puerto Plata, Tubagua, 25-iii-1970, E. Marcano,
#6196 (IIBZ); 1 ♂, Prov. Santo Domingo, Los Minas, vi-1971, E. Marcano & E. Ortega, #9706 (IIBZ); 1 ♂, Prov.
San José de Ocoa, 10-iv-1976, E. Marcano, #19942 (IIBZ); 1 ♂, Prov. La Vega, Constanza, La Palma, pie de
Casabito, 23-xii-1958, E. Marcano & Clayton, #17 (IIBZ); 2 ♂♂, Engombe, Sto. Dgo., D.N., 4-iii-79, Auyama
(IIBZ); 1 ♀, Santo Domingo, Los Minas, 27-ix-1973, E. Marcano #14141 (IIBZ); 1 ♀, Prov. Santo Domingo, Los
Minas, vi-1971, E. Marcano & E. Ortega, #9703 (IIBZ); 1 ♀, Santo Domingo, Los Minas, 27-ix-1973, E. Marcano
#14142 (IIBZ); 1 ♀, Prov. Santo Domingo, 8-ii-1970, E. Marcano, #6014 (IIBZ); 2 ♀♀, Prov. San José de Ocoa,
La Horma, 15-v-1976, E. Marcano & A. Abud, #20142 (IIBZ); 1 ♀, Prov. Santo Domingo, Los Minas, vi-1971, E.
Marcano & E. Ortega, #9702 (IIBZ); 1 ♀, Prov. La Vega, Constanza, Yaquecillo, E. Marcano & Clayton, #31
(IIBZ); 1 ♂, 1 ♀, km 13 Autop. Las Americas, Distrito Nacional, 2-iii-1982, Claudio Caamaño (MNHNSD).
Phthiacnemia Brailovsky 2009
Diagnosis. Medium size, elongate. Body surface without metallic iridescence. Posttylar depression absent. Calli
anteriorly with two small spines, each lateral to midline, and posteriorly with two deep pits, each lateral to midline.
Anterolateral borders of pronotum dentate. Posterolateral borders smooth. Humeral angles obtuse, truncate or
gently subacute. Corium without yellowish transverse fascia. Metathoracic peritreme bilobate.
Distribution. Only one species is known in the genus and this is widespread through the south of the United
States, Mexico, Central America, Cuba and South America reaching Argentina. The genus is cited for the first time
from Hispaniola.
Phthiacnemia picta (Drury)

(Fig. 12)
Cimex picta Drury, 1770: 107.
Discussion. Recognized by having the antennal segments I, II and IV black, III black with dark orange ring near
middle third; pronotum dark brown to black with anterolateral margins, humeral angles and widely transverse
fascia at posterior lobe bright orange; anterolateral margins spinated, with short subacute spine; corium black with
costal margin orange; dorsal abdominal segments black with bluish green iridescence; femora black, with basal
third and apical border reddish orange; and fore and middle tibiae black with dark yellow or dark castaneous
orange ring near middle third.
General distribution. Widespread throughout Antigua, Argentina, Barbados, Brazil, Colombia, Costa Rica,
Cuba, the Dominican Republic, El Salvador, Grenada, Guadeloupe, Guatemala, Guyana, Honduras, Mexico,
Nicaragua, Paraguay, Puerto Rico, St. Croix, Surinam, Uruguay, Venezuela and Virgin Islands. HISPANIOLA:
DOMINICAN REPUBLIC: Santo Domingo (Barber 1923a, Barber 1939, Brailovsky 2009, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♂♂, Palacio de
Engombe, Santo Domingo Oeste, 18°27.047’N 69°59.802’W, 5-viii-2011, D. Perez, S. Navarro (USNM). 2 ♀♀,
HDR-001, Parque Mirador del Norte, Santo Domingo, 18°517393’N 69°917912’W, 9-vii-2015, D. Perez, S.
Navarro (USNM). 1 ♀, Bonao, Llanos, vii-1969, J. Maldonado C. (USNM). 1 ♂, 1 ♀, Santiago Prov., Babosico,
500 m, 19°20.983’N 70°37.447’W, 12-vii-2015, D. Perez, S. Navarro (day) (USNM). 1 ♂, San Cristóbal Prov., La
Colonia to Los Cacaos, 8 km SE Los Cacaos, 650 m, 18°28.971’N 70°13.348’W, 24-vii-2017, T. Henry, M.
Guidoti, D. Perez-Gelabert (USNM). 1 ♀, San Cristóbal Prov., La Colonia to Los Cacaos, 2 km SE Los Cacaos,
550 m, 18°31.444’N 70°16.835’W, 24-vii-2017, T. Henry, M. Guidoti, D. Perez-Gelabert (USNM). 1 ♀, Ocoa
Prov., open field at the road to Los Martínez, km 3, 18°27.652’N 70°28.157’W, 600 m, M. Guidoti, D. Perez-
Gelabert (USNM); 1 ♂, Prov. María Trinidad Sánchez, Cabrera, Arroyo Salado, Baoba del Piñas, 2-xi-1975, E.
Marcano & J. Cicero, #19147 (IIBZ); 1 ♂, Prov. María Trinidad Sánchez, Cabrera, Arroyo Salado, Baoba del
Piñas, 2-xi-1975, E. Marcano & J. Cicero, #19161 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 5-iii-1976, E.
Marcano, #19671 (IIBZ); 1 ♂, Prov. Valverde, El Remolino, Río Mao, 20-viii-1975, E. Marcano, #18746 (IIBZ); 1
♂, Prov. San Cristóbal, 25-v-1969, E. Marcano, #5150 (IIBZ); 1 ♂, Prov. San Cristóbal, 29-i-1971, E. Marcano,
#8602 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 14-i-1977 E. Marcano, #21007 (IIBZ); 1 ♂, Prov. Monte
Plata, Bayaguana, Carabela, 16-x-1976 E. Marcano, #20483 (IIBZ); 1 ♂, Prov. Salcedo, Tenares, Boba Arriba, viii-
1971, E. Marcano & A. Linares, #10305 (IIBZ); 1 ♂, Prov. San Cristóbal, 29-i-1971, E. Marcano, #8611 (IIBZ); 1
♂, Prov. Barahona, Paraíso, 6-iv-1969, E. Marcano, #5046 (IIBZ); 1 ♂, Prov. Elías Piña, Carr. Internacional,
Pirámide 99, 450 m, 9-iii-1974, E. Marcano #15811 (IIBZ); 2 ♂♂, 2 ♀♀, Jarabacoa, 8-vii-79, UASD, luz (IIBZ); 1
♀, Prov. San Pedro de Macorís, Quisqueya, Ingenio Quisqueya, 26-i-1971, E. Marcano & S. Mosquete, #8574
(IIBZ); 1 ♀, Prov. San Cristóbal, vi-1971, E. Marcano, #9957 (IIBZ); 1 ♀, Prov. Monte Plata, Bayaguana,
Carabela, 16-vi-1976 E. Marcano, #20492 (IIBZ); 1 ♀, Prov. San Cristóbal, ciudad, 10-v-1976, E. Marcano,
#19985 (IIBZ); 1 ♀, Prov. Monseñor Nouel, Bonao, Jayaco—El Abanico, cruce de la carret. Duarte a Constanza,
24-iv-1970, E. Marcano, #7111. on gramíneas (IIBZ); 1 ♀, Prov. La Vega, Jarabacoa, El Pedregal, 29-vii-1967, E.
Marcano, #4265 (IIBZ); 1 ♀, Prov. Santiago, iii-1970, E. Marcano, #6874 (IIBZ); 1 ♀, Prov. La Vega, Jarabacoa,
El Pedregal, 29-viii-1967, E. Marcano, #4266 (IIBZ).
Rhytidophthia Brailovsky 2009
Diagnosis. Dorsal surface of head, calli, anterolateral borders of pronotum, scutellum, rostral segment I, acetabulae
and legs with metallic bluish purple iridescence. Posttylar depression absent. Posterior pronotal disk with low
longitudinal carinae. Humeral angles remarkably expanded into a large acute spine directed outward and forward.
Calli with two large projections, each lateral to midline (shorter in females). Scutellum wider than long. Femora
ventrally armed. Hind femora dorsally tuberculate. Male hind tibiae ventrally armed. Metathoracic peritreme
bilobate. Abdominal sterna with medial furrow.
Distribution. Previously known only from Cuba. Only one species is included.
Rhytidophthia splendida (Valdes)

(Fig. 13)
Phthia splendida Valdes, 1910: 431.
Discussion. Atractive species with corium black with widely yellowish orange transverse fascia; connexivum
yellowish orange with posterior third black; dorsal abdominal segments bright orange; and abdominal sterna III–VI
bright yellowish orange with black discoidal spot at each side of midline.
General distribution. Cuba (Valdes 1910, Brailovsky 2009, Packauskas 2010). This species is cited for the
first time from Hispaniola.
Material examined. New records. HISPANIOLA. HAITI: 1♀, Port Prince, 23–26-ii-1908, Dr. M. Cameron
(BMNH).
Tribe Chariesterini
Chariesterus De Laporte 1832
Diagnosis. Slender, elongate. Head subquadrate, deeply cleft medially with a deep pit in front of each ocellus.
Antennal segment I three sided. Antennnal segment III flattened and rounded into a leaf-like plate distally.
Antenniferous tubercles armed with the spine projecting forward. Juga and tylus strongly and abruptly deflexed.
Humeral angles with prominent acute spine directed outward. Mesosternun and metasternum deeply sulcate.
Distribution. The genus contains 12 species and occurs throughout Canada, United States, Mexico, Central
America, Greater and Lesser Antilles, and South America, reaching Argentina (Ruckes 1955, Packauskas 2010). In
this contribution the genus is cited for the first time from Hispaniola.
Chariesternus gracilicornis Stål

(Fig. 14)
Chariesternus gracilicornis Stål, 1870: 178.
Discussion. This species has the metathoracic peritreme with the auricle simple, not bilobed; dilation of antennal
segment III narrowly elliptical in outline; postocular tubercle exposed; rostrum reaching posterior border of
mesosternum; anterolateral margins of pronotum with irregularly small denticles; humeral angles unispinose, with
the spine large, acute, and directed outward; femora distally armed; and abdominal sterna without rows of
subcircular black dots.
General distribution. This species is limited to some of the West Indies. Antigua, Cuba, the Dominican
Republic, Guadeloupe, Jamaica, Puerto Rico, St. Croix, St. Eustatius and Virgin Islands. HISPANIOLA.
DOMINICAN REPUBLIC: Santo Domingo (Stål 1870, Fracker 1919, Barber 1923a, b, Barber 1939, Ruckes
1955, Zayas 1988, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 4 ♂♂, 2 ♀♀, HDR-021,
Sendero Rabo de Gato, Parque Nacional Sierra de Bahoruco, Independencia Prov., 415 m, 18°19.042’N
71°34.279’W, 31-viii-2015, D. Perez, R. Bastardo, A. Sánchez. Day + UV light (USNM). 1 ♂, Samaná, La
Majagua, 1-xii-1993, J. B. Heppner (FSCA). 1 ♂, 1 ♀, La Romana prov., 7.5 km N Guaymate, 15-v-1992, R.
Turnbow (UGAG). 1 ♀, La Estrelleta, 10 km E. Hondo Valle, 7-viii-1979, C. W. O’Brien (USNM). 1 ♀, S. C., 2
kms Najayo, 5-viii-1970, G. B. Marshall (UNAM). 1 ♂, 1 ♀, Azua, 8 km NE Padre Las Casas, Río Las Cuevas,
580 m, 18°46’N 70°53’W, 3–4-x-1991, C. Young, S. Thompson, R. Davidson, J. Rawlins (CMNH). Biological
note: In arid thornscrub. 1 ♀, Prov. San Juan, 28-xii-1979, Marcano y Reynoso (MNHNSD). 1 ♀, Distrito
Nacional, La Malena de Boca Chica, 20-ix-1919, Mata y Dguez (MNHNSD). 1 ♂, 1 ♀, Distrito Nacional,
Engombe, 20-ix-1979, Reynoso (MNHNSD). 1 ♂, Puerto Plata Prov., La Cumbre, 26-xii-1978, Domínguez, Silfa
(MNHNSD). 1 ♂, Samaná Prov., Samaná, Pto. Francés Viejo, 12-iv-1979, Domínguez (MNHNSD). 1 ♀, Prov.
Santiago, Licey al Medio, Tamboril, 19-ix-1979, Marcano (MNHNSD). 1 ♂, 1 ♀, Prov. Elias Piña, Pedro,
Guayajayuco, 9-xi-80, Marcano y Cicero (MNHNSD). 1 ♀, Prov. La Vega, 24-vi-70, Domínguez (MNHNSD). 1
♀, Dominican Republic, 10-xii-69, Domínguez (MNHNSD). 1 ♂, Distrito Nacional, Santo Domingo, Sans Souci,
21-vii-80, Marcano (MNHNSD). 1 ♂, Prov. Salcedo, Salcedo, Las Curvas, 29-vii-1979, Reynoso (MNHNSD). 2
♂♂, 1 ♀, Pedernales, 2 km SE Los Tres Charcos, 5 km NW Oviedo, 17 50’N 71 27’W, 35 m, 15-vii-92, R.
Davidson, J. Rawlins, S. Thompson, C. Young (CMNH). Biological note: roadside vegetation. 1 ♂, 9 ♀♀, Distrito
Nal., Santo Domingo, Parque Paseo de los Indios, 18°26’53”N 69°56’39”W, 60 m, 1-10-xi-2002, W. A. Zanol
(CMNH). 2 ♂♂, La Altagracia, 7 km NNE Boca de Chavon at Rio Chavon, 18°28’N 68°52’W, 20 m, 3-vii-1992, J.
Rawlins, S. Thompson, C. Young, R. Davidson (UNAM). Biological note: Riparian vegetation. 1 ♂, Barahona, 9.2
km NW Paraiso, confluence of Rio Nizao and Rio Coltico, 18°03’N 71°12’W, 230 m, 9-10-viii-1990, J. Rawlins,
S. Thompson (CMNH). 1 ♀, San Juan, 6 km W San Juan, 7-viii-79, L. B. O’Brien (UNAM). 2 ♀♀, La Altag., 4
km NW Boca de Yuma, 1-viii-79, C. W. O’Brien (UNAM). 6 ♂♂, 3 ♀♀, La Altagracia, 2 km SW Campo Nuevo,
1-viii-79, G. B. Marshall (UNAM). 1 ♂, San Pedro, 7 km W San Pedro de Macorís, 31-vii-79, G. B. Marshall
(UNAM). 1 ♂, 1 ♀, Peravia, 13 km NW Bani, 6-viii-79, G. B. Marshall (UNAM). 1 ♂, 5 ♀♀, San Cristóbal, 2 km
S Najayo, 5-viii-79, G. B. Marshall (UNAM). 1 ♂, 1 ♀, La Vega, 30 km E El Rio, 3-viii-79, L. B. O’Brien
(UNAM). 2 ♂♂, 3 ♀♀, San Cristóbal, 16-viii-76, H. Ludovino Domínguez, 8-x-81 (UNAM). 1 ♂, 1 ♀, La
Altagracia, 31 km N Higuey, 1-viii-79, L. B. O’Brien (UNAM). 1 ♂, San Cristóbal, 8-ii-81, H. Ludovino, 2 ♂♂, 7
♀♀, 24-i-76 (UNAM). 1 ♀, Peravia, 9 km E. San José de Ocoa, 8-viii-79, C. W. O’Brien, G. B., Marshall
(UNAM). 1 ♀, Sánchez, 7-12-vi-1915 (UNAM). 1 ♂, Prov. Barahona, 5 km NW Barahona, Agr. Exp. Sta., 29-30-
iv-78, G. B. Fairchild, R. E. Woodruff, noct. (UNAM). 5 ♂♂, 4 ♀♀, Monseñor Nouel Prov., Ingenio, old field nr.
Duarte Rd., km 76, 18°52.762’N 70°21.996’W, 200 m, 4.viii.2017, M. Guidoti, D. Perez-Gelabert (USNM). 1 ♂, 2
♀♀, San Pedro de Macoris Prov., Juan Dolio, forest on north side of rd., 18°25.165’N 69°28.059’W, nr sea level, 2-
viii-2016, D. Perez, S. Navarro, J. Borbón (USNM). 3 ♂♂, Azua Prov., old field at North edge of Pueblo Viejo,
18°24.455’N 70°45.796’W, 70 m, 2-viii-2017, T. Henry, M. Guidoti, D. Perez-Gelabert (USNM). 3 ♂♂, 1 ♀, San
Cristóbal Prov., La Sombra, 5 km NE of Los Cacaos, 18° 32.229’N 70°20.966’W, 700 m, 24-vii- 2017, T. Henry,
M. Guidoti & D. Perez-Gelabert (USNM). 2 ♀♀, Trail to Monte Diego de Ocampo, Santiago Prov., 750-800 m, 10-
vii-2015, 19°34.326’N 70°43.786’W, D. Perez, S. Navarro. Day + UV light (USNM). 1 ♂, San Cristóbal Prov., La
Colonia to Los Cacaos, 8 km SE Los Cacaos, 650 m, 18°28.971’N 70°13.348’W, 24-vii-2017, T. Henry, M.
Guidoti & D. Perez-Gelabert (USNM). 1 ♂, Ocoa Prov., open field at the road to Los Martínez, km 3, 18°27.652’N
70°28.157’W, 600 m, M. Guidoti, D. Perez-Gelabert (USNM). 1 ♀, Santo Domingo Oeste, km 38 Duarte Rd.,
18°38.404’N 70°09.765’W, 17-viii-2016, D. Perez, S. Navarro, J. Borbón (USNM). 1 ♀, Monseñor Nouel Prov.,
Bonao, old field with gate inside town, 18°55.624’N 70°25.701’W, 250 m, M. Guidoti, D. Perez-Gelabert
(USNM). 1 ♂, 1 ♀, DR-2016-004 (USNM). 5 ♂♂, 3 ♀♀, Ciudad Trujillo, xi-1950, N.L.H. Krauss (USNM). 3 ♂♂,
1 ♀, 6 km W. of Ciudad Trujillo, R. D., 16-20-xii-1955, J. Maldonado-Capriles, Coll. (USNM). 1 ♀, Santo
Domingo, 7.8.05, WI, viii, Busck (USNM). 1 ♀, Duarte, Santo Domingo City, 21-vii-1917, (358), Harold Morrison
(USNM). 1 ♀, La Estrel., 10 km E. Hondo Valle, 7-viii-1979, C. W. O’Brien (USNM). 1 ♀, Bonao, llanos, vii-
1969, J. Maldonado C. (USNM). 7 ♂♂, 1 ♀, 6 km W. of Ciudad Trujillo, R. D., 19-20-xii-1955, J. Maldonado-
Capriles (USNM). 1 ♂, 1 ♀, Seashore W. of Ciudad Trujillo, R. D., 21-xii-1955, J. Maldonado-Capriles. (USNM).
1 ♂, San Juan, R. D., 1-30-vii-1955, Rodriguez, Paniagua (USNM). 1 ♂, 1 ♀, Peravia, 2 km. E. Nizao, Rio Nizao,
5-viii-1979, O’Brien & Marshall (USNM); 5 ♂♂, Prov. San Cristóbal, Campus Loyola, 29-i-1971, E. Marcano,
#8684 (IIBZ); 4 ♂♂, Prov. Barahona, Playa Saladilla, 9-vii-1966, E. Marcano, #3319 (IIBZ); 1 ♂, Prov. Barahona,
Playa Saladilla, 9-vii-1966, E. Marcano, #3320 (IIBZ); 1 ♂, Prov. San Cristóbal, 29-i-1971, E. Marcano, #8626
(IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 29-i-1971, E. Marcano, #8687 (IIBZ); 1 ♂, Prov. Santo
Domingo, 8-II-1970, E. Marcano, #6018 (IIBZ); 1 ♂, Prov. Monte Plata, Bayaguana, Carabela, 16-x-1976, E.
Marcano, #20485 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8111 (IIBZ); 1 ♂,
Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8098 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus
Loyola, 8-x-1970, E. Marcano, #8103 (IIBZ); 1 ♂, Prov. Hato Mayor, Guamira, km 15, 21-xi-1965, E. Marcano,
#2739 (IIBZ); 1 ♂, Prov. Monte Plata, Bayaguana, Carabela, 16-x-1976 E. Marcano, #20484 (IIBZ); 1 ♀, Prov.
Barahona, Playa Saladilla, 9-vii-1966, E. Marcano, #3319 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 8-x-
1970, E. Marcano, #8115 (IIBZ); 1 ♀, Santo Domingo, Los Minas, 1-ix-1973, E. Marcano #14125 (IIBZ); 1 ♀,
Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8099 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus
Loyola, 8-x-1970, E. Marcano, #8102 (IIBZ); 1 ♀, Prov. Santo Domingo, Engombe, 20-xi-1970, E. Marcano & A.
Abud, #11624 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8110 (IIBZ); 1 ♀, Prov.
Monte Cristi, Villa Vázquez, Villa Copa, 5-viii-1965, E. Marcano, #1831 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus
Loyola, 8-x-1970, E. Marcano, #8096 (IIBZ); 1 ♀, Prov. Pedernales, entre Oviedo y Pedernales, 24-iii-1967, E.
Marcano, #4436 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8097 (IIBZ); 1 ♀,
Prov. Monte Plata, Bayaguana, Carabela, 16-x-1976 E. Marcano, #20482 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus
Loyola, 8-x-1970, E. Marcano, #8112 (IIBZ); 1 ♀, Prov. Santo Domingo, Villa Mella, Sabana de San Felipe, 22-xi-
1965, E. Marcano, #2741 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8100; 1 ♂,
Prov. María Trinidad Sánchez, Cabrera, Arroyo Salado, Baoba del Piñas, 2-xi-1975, E. Marcano & J. Cicero,
#19159 (IIBZ); 2 ♂♂, Prov. Santo Domingo, El Cachón de la Rubia, 16-x-1965, E. Marcano, #2521 (IIBZ); 1 ♂,
Prov. Dajabón, Loma de Cabrera, La Garrapata, 20-iv-1973, E. Marcano #13892 (IIBZ); 1 ♂, Prov. San Cristóbal,
Campus Loyola, 10-x-1970, E. Marcano #8152 (IIBZ); 1 ♂, Prov. Dajabón, Restauración, Villa Anacaona, Neita,
22-viii-1975, E. Marcano & J. Cicero, #18905 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 16-xii-1971, E.
Marcano, #10502 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 9-vi-1972, E. Marcano, #12400 (IIBZ); 1 ♂,
Prov. María Trinidad Sánchez, Nagua, El Pozo, 2-xi-1975, E. Marcano, #19176 (IIBZ); Prov. Santo Domingo,
Guerra, La Joya, 27-iv-1972, E. Marcano & R. Hansen, #12127 (IIBZ); 1 ♂, Prov. La Altagracia, Boca de Yuma,
junto a la boca, 3-xii-1976, E. Marcano, #20898 (IIBZ); 1 ♂, Santo Domingo, Engombe, 4-viii-1977, E. Marcano,
#21799 en maíz (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 16-xii-1971, E. Marcano, #10487; 1 ♀, Prov. El
Seibo, El Morro de Miches, 19-viii-1972, E. Marcano, #12752 (IIBZ); 1 ♀, Prov. San Cristóbal, 29-i-1971, E.
Marcano, #8618 (IIBZ); 1 ♀, Prov. Pedernales, Olivares, prox. Pedernales, 26-vi-1966, E. Marcano, #4452. on
maíz (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 5-xii-1970, E. Marcano, #8279 (IIBZ); 1 ♀, Prov. Santiago,
Rincón de Piedra, El Carrizal entre los ríos Lodoso y Bajamillo, 1-i-1978, E. Marcano, #21848 (IIBZ); 1 ♀, Prov.
San Cristóbal, Campus Loyola, 10-xi-1970, E. Marcano, #8239 (IIBZ); 1 ♀, Prov. Hato Mayor, Yamasá, El
Reparadero, Arroyo Aguas Sucias, 6-iii-1971, E. Marcano & H. Marión, #9235 (IIBZ); 1 ♂, Prov. Elías Piña,
Pedro Santana, Guayayuco, Carret. Internacional, 22-viii-1975, E. Marcano & J. Cicero, #18910 (IIBZ); 1 ♂, Prov.
Azua, Padre de las Casas, i-1972, E. Marcano, #10952 (IIBZ); 1 ♂, Prov. San Cristóbal, 10-xi-1970, E. Marcano,
#8238 (IIBZ); 1 ♂, Prov. Santiago Rodríguez, Sabana de La Caoba, La Caoba, 5-v-1972, E. Marcano & A. Abud,
#11427 (IIBZ); 1 ♂, Prov. San Cristóbal, 10-xi-1970, E. Marcano, #8228 (IIBZ); 1 ♂, Prov. San Cristóbal, 10-xi-
1970, E. Marcano, #8229 (IIBZ); 2 ♂♂, Prov. El Seibo, Miches, El Morro, 1-iv-1973, E. Marcano #13254 (IIBZ);
1 ♂, Prov. Peravia, Baní, La Baría, 27-ii-1975, E. Marcano & J. Cicero, #18066 (IIBZ); 1 ♂, Prov. San Cristóbal,
10-xii-1970, E. Marcano, #8219 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 8-x-1970, E. Marcano, #8048
(IIBZ); 1 ♂, Prov. San Cristóbal, 17-xii-1970, E. Marcano & A. Abud, #6748. on batata (IIBZ); 1 ♂, Prov. San
Cristóbal, Campus Loyola, 16-xii-1971, E. Marcano, #10412 (IIBZ); 1 ♂, Prov. Santo Domingo, Guerra, La Joya,
27-iv-1972, E. Marcano & R. Hansen, #12126 (IIBZ); 1 ♂, Prov. La Romana, Boca de Chabón, 18-iv-1971, E.
Marcano, #9030 (IIBZ); 1 ♂, Prov. San Juan, Matas de Farfán, Carrera de Yeguas, 18-v-1973 E. Marcano #13962
(IIBZ); 1 ♂, Prov. San José de Ocoa, El Pinal, 10-viii-1974, E. Marcano #17067 (IIBZ); 1 ♀, Prov. San Cristóbal,
Campus Loyola, 10-xi-1970, E. Marcano, #8242 (IIBZ); 1 ♀, Prov. San Cristóbal, 10-xi-1970, E. Marcano, #8216
(IIBZ); 1 ♀, Prov. El Seibo, La Candelaria, 23-iii-1975, E. Marcano & J. Cicero, #18120 (IIBZ); 1 ♀, Prov.
Barahona, Cabral, El Firme, E. Marcano, #22278 (IIBZ); 1 ♀, Prov. San Cristóbal, 9-vi-1972, E. Marcano, #12402
(IIBZ); 1 ♀, Prov. Pedernales, Cabeza de Agua, 17-viii-1973, E. Marcano #15546 (IIBZ); 1 ♀, Prov. San Cristóbal,
Campus Loyola, 23-ii-1973, E. Marcano #13139 (IIBZ); 1 ♀, Prov. Santiago, Rincón de Piedra, El Carrizal entre
los ríos Lodoso y Bajamillo, 1-i-1978, E. Marcano, #21851 (IIBZ); 1 ♀, Prov. La Altagracia, Higuey, Macao, El
Cortecito, 10-iv-1971, E. Marcano & L. Schott, #8779 (IIBZ); 1 ♀, Prov. San Cristóbal, 30-iv-1976, E. Marcano,
#19959 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 25-iv-1975, E. Marcano, #18269 (IIBZ); 1 ♀, Prov. San
Cristóbal, 8-v-1973, E. Marcano #13569 (IIBZ); 1 ♀, Prov. Azua, Padre Las Casas, río Las Cuevas, 28-iv-1973 E.
Marcano #13529 (IIBZ); 1 ♀, Prov. La Altagracia, Higuey, Macao, El Cortecito, 10-iv-1971, E. Marcano & L.
Schott, #8800; 2 ♂♂, Prov. La Vega, Rincón a Jima, 24-iv-1970, E. Marcano & R. Hansen, #7154 (IIBZ); 2 ♂♂,
Prov. Santo Domingo, Engombe, 9-v-1972, E. Marcano & A. Abud., #12150 (IIBZ); 1 ♂, 1 ♀, Prov. Santo
Domingo, Engombe, 18-xii-1969, E. Marcano & A. Abud, #5699 (IIBZ); 1 ♂, 1 ♀, Prov. El Seibo, El Morro de
Miches, 19-viii-1972, E. Marcano, #12748 (IIBZ); 1 ♂, 1 ♀, Prov. Santo Domingo, Guerra, La Joya, 27-iv-1972,
E. Marcano & R. Hansen, #12125 (IIBZ); 2 ♂♂, Prov. La Altagracia, Higuey, Macao, El Cortecito, 10-iv-1971, E.
Marcano & L. Schott, #8799 (IIBZ); 1 ♂, 1 ♀, Prov. Pedernales, entre Oviedo y Pedernales, 9-xi-1974, E. Marcano
#17404 (IIBZ); 1 ♂, Prov. La Vega, Rincón a Jima, 24-iv-1970, E. Marcano & R. Hansen, #7154 (IIBZ); 2 ♂♂,
Prov. Azua, Cortés, 27-iii-1970, E. Marcano, #6529. on Capparis spinosa (IIBZ); 1 ♂, 1 ♀, Prov. Santo Domingo,
Laguna de Guerra (Los Benjamines), 24-x-1971, E. Marcano & J. Cicero, #19117 (IIBZ); 1 ♂, Prov. San Cristóbal,
10-xi-1970, E. Marcano, #8230 (IIBZ); 1 ♂, 1 ♀, Prov. María Trinidad Sánchez, Cabrera, Arroyo Salado, Baoba
del Piñas, 2-xi-1975, E. Marcano & J. Cicero, #19160 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 10-xi-
1970, E. Marcano, #8251 (IIBZ); 1 ♂, Prov. Valverde, Mao, 4-vi-1973, E. Marcano #13603 (IIBZ); 2 ♀♀, Prov.
Pedernales, Cabeza de Agua, 17-viii-1973, E. Marcano #15521 (IIBZ); 1 ♀, Prov. San Juan, El Cercado, Jorgillo,
22-vii-1973 E. Marcano #14339 (IIBZ); 1 ♀, Prov. San Pedro de Macorís, Boca de Cumayasa, 26-i-1973, E.
Marcano #13088 (IIBZ); 1 ♀, Prov. Santiago, San José de las Matas, Los Montones Arriba, 16-iv-1973, E.
Marcano #13647 (IIBZ). HAITI: 1 ♂, Camp Perrin, 1000 ft., 8-x-1927 (UNAM). 2 ♂♂, 2 ♀♀, Port au Prince, i-ii,
P. R. Uhler (USNM). 1 ♀, Fond-des-Negres, 12-vi-1930, H. L. Dozier (USNM).
Tribe Hypselonotini
Key to Hispaniola Genera of Hypselonotini
1. Hind femora armed ventrally with two or more spines. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

- Hind femora unarmed.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
2. Hind femora clavate, swollen and bearing numerous ventral spines on distal half; humeral angles produced into long, sharp,
outwardly directed spine; body length less than 10 mm . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Zicca Amyot and Serville
- Hind femora armed ventrally with two subdistal spines and one row of small spines or tubercles; humeral angles obtuse or
truncate, never spined; body length generally longer than 12 mm . . . . . . . . . . . . . . . . . . . . . Anasa Amyot and Serville (partly)
3. Tylus elevated and compressed above juga to form a carina between antennal bases; veins of hemelytral membrane numerous,
irregularly anastomosing, or almost reticulated . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
- Tylus not compressed and elevated above juga; veins of hemelytral membrane less numerous, simple or at most slightly
branched or furcate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
4. Antennal segments II and III compressed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .Sethenira Spinola … S. ferruginea Stål
- Antennal segments II and III cylindrical. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Althos Kirkaldy … A. obscurator (Fabricius)
5. Shelf-like plate present beneath antenniferous tubercle extending over lower rim of antennal articulation . . . .Catorhintha Stål
- Head below antenniferous tubercle lacking a raised shelf-like plate . . . . . . . . . . . . . . . . . . . Anasa Amyot and Serville (partly)
Althos Kirkaldy 1904
Diagnosis. Medium sized bugs usually under 12 mm, oblong and dull colored. Hemelytral membrane with
numerous anastomosing veins. Tylus elevated and compressed. Antennal segments II–III cylindrical. Pronotum
nearly flat, anterolateral angles sharply carinate, acutely produced anteriorly into a short tooth. Legs slender.
Abdomen not swollen. Femora unarmed. Female genital plates laminate, and elongate.
Distribution. The genus includes 24 species, and these are abundant and diverse in the New World from
United States to Northern Argentina and Chile, and Greater and Lesser Antilles (Brailovsky 1990, Packauskas
2010). In this contribution the genus is cited for the first time from Hispaniola.
Althos obscurator (Fabricius)

(Fig. 15)
Coreus obscurator Fabricius, 1803: 200.
Discussion. Recognized by having antenniferous tubercle armed exteriorly with short spine directed forward;
humeral angles rounded and not raised or exposed; anterior and posterior lobe of metathoracic peritreme yellow;
legs yellow with numerous small black dots; connexival segments bicolorous alternating yellow and black; dorsal
punctures not deeply excavated; and gonocoxa I short, not reaching the middle third of paratergite IX.
General distribution. Widely distributed throughout Argentina, Brazil, British Guiana, Colombia, Costa Rica,
Cuba, Dominica, Grenada, Guadeloupe, Guatemala, Mexico, Panama, Paraguay, Peru, Puerto Rico, Surinam,
Trinidad, Uruguay, the United States and Venezuela (Barber 1939, Brailovsky 1986, Packauskas 2010). This
species is cited for the first time from Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♂♂, Rd. to Monteada
Nueva, N. of Polo, Barahona Prov., 18°06.985’N 71°15.246’W, 810 m, 28-viii-2014, D. Perez, C. de Soto M.
(USNM). 1 ♂, HDR-002 Trail to Monte Diego de Ocampo, Santiago Prov., 750–800 m, 19°34.326’N
70°43.786’W, 10-vii-2015, D. Perez, S. Navarro. Day + UV (USNM). 1 ♂, Trail to Los Guineos, Polo, Barahona
Prov., 18°08.071’N 71°15.701’W, 16-viii-2011, D. Perez, S. Navarro, R. Gomez (day) (USNM). 1 ♀, Rd. to
Monteada Nueva, N. of Polo, Barahona Prov., 18°06.985’N 71°15.246’W, 810 m, 28-viii-2014, D. Perez, C. de
Soto M. (USNM). 2 ♂♂, 3 ♀♀, Prov. El Seybo, Sabana de la Mar, Alto de la Piedra, 29-iv-1980, Marcano—Mota
(MNHNSD). 1 ♂, 1 ♀, Prov. Puerto Plata, La Cumbre, 28-xii-1978, Domínguez, Silfa (MNHNSD). 1 ♀, Prov.
Peravia, Arroyo Cañas, 5-viii-1980, A. Norrbom (CMNH). 1 ♀, Prov. La Altagracia, Higuey, Verón, 10-viii-1979,
Marcano, Mota, Reynoso (MNHNSD). 9 ♂♂, 1 ♀, La Vega Prov., 4 km E. of La Ciénaga, Manabao, 3050 ft.,
19°04’47”N 70°49’29”W, 4–8-April-2000, T. J. Henry & R. E. Woodruff (USNM). 1 ♂, La Vega Prov.: Parque
Nacional Armando Bermúdez, La Ciénaga, Manabao, 3050 ft., 19°03’45”N 70°51’50”W, 19-April-2000, T. J.
Henry & R. E. Woodruff (USNM); 1 ♂, Prov. Salcedo, Gaspar Hernández, Blanco, 12-V-1973, E. Marcano #14685
(IIBZ); 1 ♂, Prov. Puerto Plata, La Cumbre, 3-VIII-1974, E. Marcano #17196 (IIBZ); 1 ♂, Prov. Salcedo, Gaspar
Hernández, Blanco, 12-V-1973, E. Marcano #14673 (IIBZ); 1 ♂, Prov. Azua, El Número, Diciembre 1970, E.
Marcano, #9367 (IIBZ); 1 ♂, 1 ♀, Prov. Valverde, Loma de Guayacanes-El Mamey, 16-VII-1972, E. Marcano
#12906 (IIBZ); 1 ♂, 1 ♀, Prov. Santiago, Los Ramones, prox. a Palo Alto, Cordillera Septentrional, 1000m, 1-VIII-
1965, E. Marcano, #1813 (IIBZ); 7 ♂♂, Prov. Santiago, Los Ramones, prox. a Palo Alto, Cordillera Septentrional,
1000m, 1-VIII-1965, E. Marcano, #1812 (IIBZ); 3 ♂♂, Prov. Barahona, Polo, Monteada Nueva, 25-III-1967, E.
Marcano, #4008 (IIBZ); 1 ♂, Prov. Pedernales, Agua Negra, 22-III-1967, E. Marcano, #3899 (IIBZ); 1 ♂, 2 ♀♀,
Prov. La Vega, Constanza, La Palma, pie de Casabito, 23-XII-1958, E. Marcano & Clayton, #12; 2 ♀♀, Prov.
Santiago, Los Ramones, prox. a Palo Alto, Cordillera Septentrional, 1000m, 1-VIII-1965, E. Marcano, #1812
(IIBZ); 1 ♀, Prov. Pedernales, Colonia La Altagracia, 22-III-1967, E. Marcano, #3930 (IIBZ); 2 ♂♂, Prov. Hato
Mayor, Yamasá, El Reparadero, Arroyo Aguas Sucias, 6-III-1971, E. Marcano & H. Marión, #9237 (IIBZ); 1 ♂,
Prov. Monseñor Nouel, Bonao, Arroyo Blanco, vía Casabito a 3 km de El Abanico, 24-III-1970, E. Marcano & G.
Maldonado, #6229 (IIBZ); 2 ♂♂, Prov. Puerto Plata, La Cumbre, 3-VIII-1974, E. Marcano #17145 (IIBZ); 2 ♂♂,
1 ♀, Prov. Puerto Plata, La Cumbre, 3-VIII-1974, E. Marcano #17144 (IIBZ); 3 ♂♂, 1 ♀, Prov. Puerto Plata, La
Cumbre, 3-VIII-1974, E. Marcano #17134 (IIBZ); 1 ♂, Prov. Puerto Plata, La Cumbre, 3-VIII-1974, E. Marcano
#17176 (IIBZ); 1 ♂, 1 ♀, Prov. Monseñor Nouel, Bonao, Arroyo Blanco, vía Casabito a 3 km de El Abanico, 24-
III-1970, E. Marcano & G. Maldonado, #6250 (IIBZ); 1 ♂, Prov. Barahona, Paraiso, Leonardo, 29.III.1972, E.
Marcano & J. Fernández, #11328 (IIBZ); 3 ♂♂, 1 ♀, Prov. Monseñor Nouel, Bonao, Arroyo Blanco, vía Casabito
a 3 km de El Abanico, 24-III-1970, E. Marcano & G. Maldonado, #6218 (IIBZ); 1 ♂, 2 ♀♀, Prov. Hato Mayor,
Yamasá, El Reparadero, Arroyo Aguas Sucias, 6-III-1971, E. Marcano & H. Marión, #9234 (IIBZ); 1 ♀, Prov.
Hato Mayor, Yamasá, El Reparadero, Arroyo Aguas Sucias, 6-III-1971, E. Marcano & H. Marión, #9236 (IIBZ); 1
♀, Prov. El Seibo, Miches, El Morro, 1-IV-1973, E. Marcano #13250 (IIBZ); 1 ♀, Prov. Monseñor Nouel, Bonao,
Arroyo Blanco, vía Casabito a 3 km de El Abanico, 24-III-1970, E. Marcano & G. Maldonado, #6229 (IIBZ).
Anasa Amyot and Serville 1843
Diagnosis. Medium sized, robust, elongate oval; head subquadrate, not strongly declivent, produced noticeably
beyond bases of antennae; tylus not compressed and raised above jugum; antenniferous tubercles unarmed or
armed; postocular tubercle usually swollen; rostrum reaching or extending beyond the middle third of
mesosternum; head below the antenniferous tubercle lacking a raised shelf-like plate; pronotum strongly declivent;
anterolateral margins smooth or serrate, or nodulose, markedly narrowing anteriorly; humeral angles obtuse or
laterally expanded and directed outward and forward, never acutely spined; hind femora slender, unarmed or armed
ventrally with one or more subdistal spines; connexivum exposed.
Distribution. A large genus with 80 species, widely distributed in the Nearctic and Neotropical regions
(Brailovsky 1985, Brailovsky 2001, Brailovsky 2008, Brailovsky 2106, Brailovsky & Barrera 2008, Brailovsky &
Barrera 2009a, Packauskas 2010, Brailovsky 2017). Five species occur in Hispaniola.
Key to Hispaniola Species of Anasa
1. Hind femora armed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . A. scorbutica (Fabricius) (partly)

- Hind femora unarmed or at most with some nodular processes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
2. Antenniferous tubercle unarmed or at most with some nodular processes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
- Antenniferous tubercle armed with short to large spine . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
3. Humeral angles expanded sideways and projecting in a sharp process; antennal segment I long, cylindrical and the apex
slightly enlarged; anterior and posterior lobe of metathoracic scent gland peritreme black to dark brown; tylus short, giving to
the head the appearance of being quadrangular . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . A. acutangula Stål
- Humeral angles not prominent and the apex obtusely rounded; antennal segment I short, thick and flared on the inside face;
anterior and posterior lobe of metathoracic scent gland peritreme yellow; tylus long, giving to the apex of head a triangular
appearance . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .A. tristis (De Geer)
4. Antenniferous tubercle armed with a long spine, the length of which is equivalent to one third of the total length of antennal
segment I . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . A. scorbutica (Fabricius) (partly)
- Antenniferous tubercle armed with short to medium sized spine, always shorter than one third of the total length of antennal
segment I . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
5. Antennal segments II and III distinctly annulate, alternating yellow and black rings . . . . . . . . . . . . . . . . A. bellator (Fabricius)
- Antennal segments II and III chestnut orange scattered with reddish brown tubercles . . . . . . . . . . . . . . . . A. pisina Brailovsky
Anasa acutangula Stål

(Fig. 16)
Anasa acutangula Stål, 1870: 196–197.
Discussion. The following characters differentiate this species from the remaining members of the genus Anasa
present in Hispaniola: humeral angles expanded sideways and projecting in sharp process; antennal segment I long,
cylindrical and apically slightly enlarged; antenniferous tubercles unarmed; head dorsally ochre or yellowish with
two black, complete, longitudinal stripes that run beside the median line; anterior and posterior lobe of
metathoracic peritreme black; frontal angles of the pronotum obtuse, never touching the cephalic border; and tylus
short, giving to the head the appearance of being quadrangular.
General distribution. Previous to this paper only known from Cuba (Brailovsky 1985, Packauskas 2010).
This species is cited for the first time from Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, Puerto Plata, Reserva
Científica Isabel de Torres, near headqrs., 112 m, 19°46’42.3”N 70°42’13.4”W, 21-vi-2010, S. W. Lingafelter
(USNM). 1 ♂, Masacó, S. of Polo, Barahona Prov., 18°02.495’W 71°18.408’W, 812 m, uv light, 28-viii-2014, D.
Perez, C. de Soto M. (USNM). 1 ♀, DR-2016-036, La Vega Prov., Arroyazo, Reserva Científica Ebano Verde, day
and night collecting with UV light, 19°02.327’N 70°32.336’W, 1154 m, 27-viii-2016, D. Perez, R. Bastardo
(UNAM).
Anasa bellator (Fabricius)

(Fig. 17)
Cimex bellator Fabricius 1787: 286.
Discussion. A unique set of characters separate this species: antennal segments II and III yellow with two wide
black rings, one subbasal and the other subdistal; antennal segment IV black with apical third yellow; antenniferous
tubercles armed with short acute spine that reaches base of antennal segment I; femora unarmed; pronotum and
scutellum pale chestnut orange; humeral angles of pronotum slightly exposed but not extended forward; and
abdominal sternites and genital plates pale chestnut orange.
General distribution. Widely distributed throughout Mexico, Central America, West Indies and South
America: Belize, Bolivia, Brazil, British Guiana, Colombia, Costa Rica, Cuba, Dominica, the Dominican Republic,
Ecuador, El Salvador, French Guiana, Grenada, Guadaloupe, Guatemala, Honduras, Mexico, Nicaragua, Panama,
Peru, St. Vincent, Suriname, Trinidad, and Venezuela. HISPANIOLA: DOMINICAN REPUBLIC: La Plata, and 21
km N of Higuey (Brailovsky 1985, Brailovsky 2001, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, 2 ♀♀, DR-2016-007,
San Cristóbal Prov., On dirt road going up, Rancho Viejo, 18°28.332’N 70°15.553’W, 950 m, 10-viii-2016, D.
Perez, C. de Soto Molinari, D. Pou (USNM). 1 ♀, San Cristóbal Prov., La Colonia to Los Cacaos, 8 km SE Los
Cacaos, 650 m, 18°28.971’N 70°13.348’W, 24-vii-2017, T. Henry, M. Guidoti & D. Perez-Gelabert (USNM). 1 ♂,
La Vega, 22 km NW Bonao, 3-ix-1997, C. W. O’Brien (UNAM). 1 ♀, La Altagracia, 21 km N. Higuey, 1-viii-79,
C. W. O’Brien, Marshall (UNAM). 1 ♀, La Plata, Hurst (UNAM). 1 ♂, La Altagracia, 31 km N Higuey, 1-viii-79,
L. O’Brien (UNAM). 11 ♂♂, 5 ♀♀, La Altagracia, Parque del Este, Caseta Guaraguao, 4.4 km SE Bayahibe,
18°19’59” N 68°48’42” 3 st., 26–27-v-2004, C. Young, J. Rawlins, J. Fetzner, C. Nuñez (CMNH). Biological note:
semihumid forest near sea, limestone. 1 ♀, Prov. El Seibo, Sabana de la Mar, Alto de la Piedra, 29-iv-80,
Domínguez, Mtz. (MNHNSD); 1 ♀, Prov. Santiago, Carretera, 19-vii-2014, A. Sánchez (IIBZ).
Anasa pisina Brailovsky
(Fig. 18)
Anasa pisina Brailovsky, 2001, 43–45.
Discussion. This species has the antenniferous tubercles armed with short, acute spine; antennal segments I–IV
chestnut orange; humeral angles exposed, produced laterally, similar to short wing and directed upward; fore and
middle femora unarmed, hind femora with blunt and inconspicuous subdistal spine-like tubercles; femora and
tibiae pale orange with black discoidal spots; and dorsal abdominal segments bright orange.
General distribution. Only known from Dominican Republic. HISPANIOLA: DOMINICAN REPUBLIC: La
Vega, vic. Salto de Aguas Blancas (Brailovsky 2001, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, Sierra Martín García,
Azua Prov., El Curro, caseta Parques, 18°22.372’N 71°01.724’W, 1,350 m, 22–23-viii-2014, D. Perez, C. Marte,
K. Rodríguez, A. Sánchez (USNM). 1 ♀, RD-218 1 km ESE Cortico Barahona prov., 1,347 m, 18°06.520’N
71°12.898’W, 9–10-iv-2004, D. Perez, B. Hierro, R. Bastardo. (d/n) (USNM). 1 ♂, 1 ♀, La Vega, vic. Salto Aguas
Blancas, 19-vii-96, R. Turnbow (UNAM); 2 ♂♂, Prov. La Vega, La Ciénaga, P. N. Armando Bermúdez,
(19°4’22”N 70°51’849”W), 1183 m, 19-viii-2006, coll. D. Perez, R. Bastardo, B. Hierro (IIBZ); 1 ♀, Prov. La
Vega, El Arroyazo, Reserva Científica Ebano Verde, 980 m, 337594 mE 2105122 mN, 4–5-vi-2005, R. Bastardo,
B. Hierro (IIBZ); 1 ♂, 2 ♀♀, Loma La Hoz, Bahoruco Oriental, 18-viii-2014, M. R. Bobadilla (IIBZ).
Anasa scorbutica (Fabricius)

(Fig. 19)
Cimex scorbuticus Fabricius, 1775: 706.
Discussion. This is probably the most common species of Anasa in Hispaniola and is characterized by having the
antenniferous tubercles armed with prominent spine, spine longer than diameter of eye, and 1/3 or longer than
antennal segment I; antennal segments II–III usually yellow to chestnut orange without black spots; all femora
ventrally armed with a sharp spine; femora and tibiae yellow with black spots; pronotal disk strongly declivent,
bearing numerous erect setigerous hairs that contrast with semidecumbent hairs of clavus and corium; anterolateral
bordes of pronotum crenate, with short spines; dorsal abdominal segments III–VI usually bright orange, and VII
black; and connexival segments III–VII black with anterior margin yellow. Body stout, moderate sized (12–14
mm).
General distribution. Wide spread from southwestern United States across Mexico, Central America, West
Indies and South America. Antigua, Argentina, Bahamas, Barbados, Belize, Bolivia, Brazil, British Guiana,
Colombia, Costa Rica, Cuba, Dominica, the Dominican Republic, Ecuador, El Salvador, French Guiana,
Galapagos, Grenada, Guadeloupe, Guatemala, Haiti, Honduras, Jamaica, Martinique, Mexico, Nicaragua, Panama,
Peru, Puerto Rico, St. Kitts, St. Vincent, Trinidad, the United States, Venezuela, and Virgin Islands. HISPANIOLA:
DOMINICAN REPUBCLIC: Santo Domingo, Ciudad Trujillo, Sánchez, Puerto Plata, La Vaca (6 km W of
Jayaco), Villa Rivas, 13 milles N of Pedro Sánchez, 7 km E of San Cristóbal, 1 km W of Boca Chica, and La Vega
(30 km E of El Rio). HAITI: St. Marc, Grand Rivière and Diquini (Barber 1923a, Brailovsky 1985, Baranowski &
Slater 1986, Brailovsky 2001, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, Santiago Prov., Trail
to Monte Diego de Ocampo, 750–800 m, 10-vii-2015, 19°34.326’N 70°43.786’W, D. Perez, S. Navarro (day+UV
light) (USNM). 1 ♂, Palacio de Engombe, Santo Domingo Oeste, 18°27.047’N 69°59.802’W, 5-viii-2011, D.
Perez, S. Navarro (USNM). 1 ♂, Monseñor Nouel Prov., rd. to Blanc o hydroelectric, 18°52.940’N 70°30.346’W,
700 m, 13-viii-2016, D. Perez, R. Bastardo, A. Sánchez. (USNM). 5 ♀♀, Palacio de Engombe, Santo Domingo
Oeste, 18°27.047’N 69°59.802’W, 5-viii-2011, D. Perez, S. Navarro (USNM). 2 ♀♀, 2 km W San Rafael del
Yuma, nr. vertedero, 75 m, 18°26.276”N 68°41.520”W, D. Perez, R. Bastardo RD-200 (USNM). 1 ♀, Cotuí, La
Piñita (EL-2T), Prov. Sánchez Ramírez, 374-464 mE 2091-355 mN, 3-viii-2003, R. H. Bastardo (USNM). Azua
Prov., old field at North edge of Pueblo Viejo, 18°24.455’N 70°45.796’W, 70 m, T. Henry, M. Guidoti, D. Perez-
Gelabert (USNM). 1 ♂, 13 km N. Pedro Sánchez, 2-viii-1979, B. Marshall (UNAM). 4 ♂♂, 3 ♀♀, 7 km E. de San
Cristóbal, 5-viii-79, Marshall (UNAM). 1 ♂, La Vega: 30 km E. El Rio, 3-viii-79, C. O’Brien (UNAM). 1 ♂, 1 ♀,
1 km W. Boca Chica, 30-v-78, O’Brien-Marshall (UNAM). 1 ♂, La Altagracia, 21 km N Higuey, 1-viii-79,
O’Brien, Marshall (UNAM). 1 ♀, Pedernales, 5 km NE Los Arroyos, 1680 m, 18-15N 71-54W, 20-x-91, J.
Rawlins, B. Davidson, C. Young, S. Thompson (UNAM). 1 ♂, 2 ♀♀, 3 km S Miches, 2-viii-79, G. B. Marshall
(UNAM). 2 ♂♂, 1 ♀, La Altagracia, 2 km SW Campo Nuevo, 1-viii-79, L. B. O’Brien (UNAM). 1 ♂, Distrito
Nacional Santo Domingo, Parque Paseo de los Indios, 18°26’53”N 69°56’39”W, 60 mt, 1-10-xi-2002, W. A. Zanol
(CMNH). 1 ♀, Peravia, Arroyo Cana, 65 mt, 7-viii-1980, A. Norrbom (CMNH). 1 ♀, Ciudad Trujillo, 23-xi-44, J.
Friauf (UNAM). 1 ♂, Santo Domingo, 10-vi-1924, D. T. Danforth (UNAM). 2 ♂♂, 1 ♂, Puerto Plata, Hurst
(UNAM). 1 ♀, Villa Riva, 19-vi-1915 (UNAM). 1 ♀, Sánchez, 22-v-1915 (UNAM). 1 ♀, La Vaca, 6 km W. de
Jayaco, 2-vi-69, Flint—Gómez (UNAM). 1 ♂, La Altagracia, 21 km N. Higuey, 1-viii-79, O’Brien—Marshall
(UNAM). 1 ♂, Prov. Puerto Plata, La Cumbre, 26.xii.1978, Domínguez, (MNHNSD). 1 ♂, Prov. La Vega,
Constanza, Valle Nuevo, 26-viii-79, Marcano (MNHNSD). 1 ♀, La Altagracia, Parque del Este, Caseta Guaraguao,
4.4 km SE Bayahibe, 18°19’59” N 68°48’42” 3 st., 26-27-v-2004, C. Young, J. Rawlins, J. Fetzner, C. Nuñez
(CMNH). Biological note: semihumid forest, near sea, limestone. 1 ♀, La Vega, Loma del Casabito, 19°03’N
70°31’W, 1380 mt, 3-xi-2002, W. A. Zanol, C. W. Young, S. Staresinic, J. Rawlins (CMNH). Biological note: Wet
cloud forest UV light. 1 ♀, Puerto Plata, La Cumbre, 26-xii-1978, Domínguez, Silfa (MNHNSD). 1 ♀, Prov. El
SeIbo, Sabana de la Mar, Alto de la Piedra, 29-iv-1980, Marcano, Aquino (MNHNSD). 1 ♀, Prov. Barahona, 4-5
km S. Barahona, 16-v-92, R. Turnbow (UGAG). 1 ♀, Pedernales, 19-20 km N Cabo Rojo, 375 mt, 10-vii-96, R.
Turnbow (UGAG). 1 ♀, La Vega, 2 km N Buena Vista, 21-vii-96, R. Turnbow (UGAG). 2 ♂♂, 5 ♀♀, Prov. La
Altagracia, Higuey, Nisibón, 28-iv-1980, Aquino, Mtz, Dguez (MNHNSD). 1 ♀, La Vaca, 6 km W. of Jayaco, 2-vi-
1969, Flint & Gomez (USNM). 1 ♂, 1 ♀, La Vega Prov., Parque Nacional Armando Bermúdez, La Ciénaga,
Manabao, 3050 ft., 19°03’45”N 70°51’50”W, 21-iv-2000, T. J. Henry & R. E. Woodruff (USNM); 1 ♂, Prov. Santo
Domingo, Engombe, 6-viii-2006, D. Perez, R. Bastardo, R. Rodríguez, A. Hilario (IIBZ); 1 ♀, Prov. Hato Mayor,
P. N. Los Haitises, Trepada Alta, 186 m, 446665 mE 2100744 mN, 7-i-2006, E. Fernández, J. A. Ottenwalder, R.
Bastardo (IIBZ); 2 ♀♀, Monseñor Nouel, Bonao, alrededor de la casa de máquinas de Hidroeléctrica de Blanco,
(18°52’50.3”N 70°29’50.5”N), 542 m, 28-x-2005, S. Vélez, R. Bastardo (IIBZ); 2 ♂♂, , Prov. Azua, El Número,
28-xii-1976, E. Marcano, #20881 en cundeamor (IIBZ); 1 ♂, 1 ♀, Prov. Azua, El Número, 28-xi-1976, E.
Marcano, #20879 en cundeamor (IIBZ); 3 ♂♂, Prov. Valverde, Esperanza, 25-vi-1965, E. Marcano, #1520 (IIBZ);
1 ♂, Prov. San Cristóbal, 29-i-1971, E. Marcano, #8604 (IIBZ); 1 ♂, Prov. San Cristóbal, 10-xi-1970, E. Marcano,
#8237 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 23.-ii-1973, E. Marcano #13133 (IIBZ); 1 ♂, Prov. San
Cristóbal, Campus Loyola, 5-iii-1976, E. Marcano, #19676 (IIBZ); 1 ♂, Prov. San Cristóbal, Catarey, El Duey, 27-
x-1968, E. Marcano & L. Schott, #4781 (IIBZ); 1 ♂, Prov. Santo Domingo, San Isidro, 20-vi-1974, E. Marcano
#16779 (IIBZ); 1 ♂, Prov. San Cristóbal, 9-vi-1972, E. Marcano, #12404 (IIBZ); 1 ♂, 1 ♀, Prov. Azua, El Número,
28-xi-1976, E. Marcano, #20880 en cundeamor (IIBZ); 1 ♂, Prov. Monte Plata, Sierra de Agua, Los Berros, 11-ix-
1977, E. Marcano, #21953 (IIBZ); 1 ♂, Engombe, Santo Domingo, D.N., 4-iii-76 (IIBZ); 1 ♀, Prov. San Cristóbal,
Campus Loyola, 1-iii-1968, E. Marcano, #4903 (IIBZ); 1 ♀, Prov. Pedernales, Banano, Río Mulito, 17-viii-1973,
E. Marcano #14096 (IIBZ); 1 ♀, Prov. La Altagracia, Higuey, i-1972, E. Marcano, #10937 (IIBZ); 1 ♀, Prov.
Monte Plata, El Horno, 22-i-1967, E. Marcano & L. Marión, #3663 (IIBZ); 1 ♀, Prov. San Cristóbal, Sainaguá, 25-
v-1971, E. Marcano & J. Familia, #10194 (IIBZ); 1 ♀, Prov. Santo Domingo, Los Minas, xi-xii-1969, E. Marcano
& E. Ortega, #5818 (IIBZ); 1 ♀, Prov. San Juan, Sabana Alta, El Guanito, 18-vi-1972, E. Marcano & R. Hansen,
#12565 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 19.-iii-1973, E. Marcano, #13218 (IIBZ); 1 ♀, Prov. San
Cristóbal, 29-i-1971, E. Marcano, #8616 (IIBZ); 1 ♀, Prov. Pedernales, Los Arroyos, 17-viii-1973, E. Marcano
#15441 (IIBZ); 1 ♀, Prov. Monte Plata, Bayaguana, Antón Sánchez, 21-x-1972, E. Marcano, #12834 (IIBZ); 1 ♀,
Prov. Santiago, Jánico, Juncalito, El Cacique, 1350 msnm, 11-x-1975, E. Marcano & J. Cicero, #19062 (IIBZ); 1
♀, Prov. San Cristóbal, Medina, 28-xii-1974, E. Marcano #17457 col. Ac. M. Velez Santana (IIBZ); 1 ♀, Prov. San
Cristóbal, Campus Loyola, 25-iv-1975, E. Marcano, #18275 (IIBZ); 1 ♀, Prov. Monte Plata, Arroyo Rancho, entre
Río Verde y Yamasá, 24-xi-1973, E. Marcano #14981 (IIBZ); 1 ♀, Prov. Barahona, Paraiso, Platón, confluencia río
Nizaito y la Cañada Cortico, 27-xi-1976, E. Marcano, #20836 (IIBZ); 1 ♀, Prov. La Vega, Reserva Científica
Ebano Verde, La Sal, 5-xii-2014, A. Sánchez, C. Esquea, P. Torres, J. A. García (IIBZ). HAITI: 2 ♂♂, 4 ♀♀, St.
Marc, W. Mann (UNAM). 2 ♂♂, Diquini, W. M. Mann (UNAM). 2 ♂♂, Grand Riviere (UNAM).
Anasa tristis (De Geer)
(Fig. 20)
Cimex tristis De Geer, 1773: 340.
Discussion. This is a robust, oblong oval species, ranging from dull yellow to dark brown and densely covered
with dark punctures; antenniferous tubercles unarmed or at most with some nodular process; antennal segments II–
III black with or without dark orange marks; head dorsally ochre with two broad, black, longitudinal stripes that
run on each side of medial line; tylus long giving the apex of head a triangular appearance; frontal angles of
pronotum projects much more than the cephalic border; anterolateral margins of pronotum yellow to ochre and
contrast with the dark coloration of pronotal disk; humeral angles obtusely rounded and not expanded;
posterolateral margins of pronotum wavy and very concave; triangular process of pronotum long and sharp; apex
of scutellum yellow; and femora unarmed.
General distribution. Throughout southern Canada, the United States, Mexico, Belize, Honduras, Cuba and
Brazil (Brailovsky 1985, Baranowski & Slater 1986, Packauskas 2010). This species is cited for the first time from
Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, La Vega Province,
Concepción de la Vega, 10–30-xii-2007, M. Micka (NMPC); 10 ♂♂, 2 ♀♀, Prov. Santo Domingo, Santo Domingo
Oeste, Haina, Finca de Engombe, 24-vii-2002, C. A. Nuñez, J. Asiático. En Cucurbita pepo. (MNHNSD); 1 ♀,
Urb. Buena Ventura, Santo Domingo, 13-iii-2017, Luis López (IIBZ).
Catorhintha Stål 1859
Diagnosis. Small to moderate size bugs (8–13 mm); dull colored grayish brown to yellowish; tylus not projecting
beyond jugae; antenniferous tubercles usually small, not occupying most of anterior head width, with or without
prominent spine; antennal segment I slightly thickened, slightly curved, one-half or more length of head; antennal
segment III usually shorter than II, and IV not flattened; humeral angles obtusely rounded, not prominent; male
hind femora not strongly incrassate; tibiae sulcate; hind tibiae not expanded, apically unarmed; connexivum
narrowly exposed.
Distribution. The genus contains 32 species and are abundant and diverse in the New World from Southern
Canada to Northern Argentina, including the Antilles. In Hispaniola two species C. guttula (Fabricius) and C.
selector Stål, were recorded (Brailovsky & Garcia 1987, Packauskas 2010, Brailovsky & Barrera 2010).
Key to Hispaniola Species of Catorhintha
1. Antenniferous tubercle unarmed; dorsal abdominal segments black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .C. selector Stål
- Antenniferous tubercle armed with a medium sized spine; dorsal abdominal segments black, scars of scent glands IV–V, and
V–VI yellow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .C. guttula (Fabricius)
Catorhintha guttula (Fabricius)

(Fig. 21)
Lygaeus guttula Fabricius, 1794: 162.

Discussion. A distinctive species with antenniferous tubercle armed exteriorly with an erect spine, directed
outward and never reaching 1/3 of antennal segment I; body size less than 9.5 mm.; femora, tibiae, thorax and
abdominal sterna dark yellow with numerous small black dots; corium with two callosities or yellow spots;
connexival segments entirely yellow or bicolorous alternating yellow and black; and inner surface of proximal half
of femora of males flattened and covered with long hairs, the same area in females flattened but only a few short
hairs present.
General distribution. Widely distributed throughout Antigua, Argentina, Bahamas, Bimini, Bolivia, Brazil,
Colombia, Cuba, the Dominican Republic, El Salvador, Ecuador, French Guiana, Grenada, Guatemala, Honduras,
Jamaica, Mexico, Nicaragua, Panama, Paraguay, Peru, Puerto Rico, St. Croix, St. John, St. Kitts, St. Lucia, St.
Thomas, Surinam, Tortola, Trinidad, the United States, Venezuela, and Virgin Islands. HISPANIOLA:
DOMINICAN REPUBLIC: Santo Domingo, San Juan, Bonao, Llanos y San Cristóbal (Barber 1923a, b, Barber
1939, Brailovsky & Garcia 1987, Packauskas 2010, Brailovsky & Barrera 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 3 ♂♂, Distrito Nacional,
Santo Domingo, Parque Paseo de los Indios, 18°36’53”N 69°56’39”W, 60 m, 1–10-xi-2002 (MNHNSD). 3 ♂♂, 1
♀, Prov. La Altagracia, Guaraguao, 29-ii–1-iii-1992, K. Guerrero, F. del Monte (MNHNSD). 1 ♂, Prov. Santiago,
Baitoa, 1-iv-1979, Mota (MNHNSD). 2 ♀♀, Prov. Azua, El Número, 22-vii-1979, Domínguez, Marcano, Reynoso
(MNHNSD). 1 ♀, Prov. Monte Cristi, Monte Cristi, 19-ix-1969, Domínguez (MNHNSD). 3 ♂♂, San Cristóbal,
16-viii-76, H. Ludovino, (UNAM). 25 ♂♂, 21 ♀♀, La Romana Centr., (338), 15-vii-1917, Harold Morrison
(USNM). 3 ♂♂, San Cristobal, 26-vii-1917 (379), Harold Morrison (USNM). 2 ♂♂, 1 ♀, Near San Pedro de
Macoris, 16-vii-1917 (339), Harold Robinson (USNM). 1 ♂, Bonao, Llanos, vii-69, J. Maldonado (UNAM). 5 ♂♂,
San Cristóbal, 16-viii-76, H. L. Domínguez, (UNAM). 4 ♀♀, San Cristóbal, 24-i-76, H. L. Domínguez (UNAM). 1
♂, San Juan, 28 km SE. San Juan, 6-viii-79, L. & C. O’Brien (UNAM); 2 ♂♂, Prov. Peravia, Baní, La Baría, 27-ii-
1975, E. Marcano & J. Cicero, #18073 (IIBZ); 2 ♂♂, Prov. La Vega, Pontón, 26-iii-1970, E. Marcano, #6403
(IIBZ); 2 ♂♂, 2 ♀♀, Prov. Azua, Las Charcas, Las Acequias, 16-i-1971, E. Marcano & A. Abud, #9111 (IIBZ); 4
♂♂, Prov. Peravia, Baní, Paya, 27-viii-1972, E. Marcano, #12625, on berenjena (IIBZ); 2 ♂♂, Prov. Peravia, Baní,
Fundación Sabana Buey, 1-vi-1972, E. Marcano & J. Cicero, #12273; 1 ♂, Prov. Peravia, Baní, Paya, 27-viii-1972,
E. Marcano, #12626, on berenjena (IIBZ); 5 ♂♂, Prov. La Vega, Pontón, 26-iii-1970, E. Marcano, #6406 (IIBZ); 2
♂♂, Prov. Peravia, Baní, Fundación Sabana Buey, 1-vi-1972, E. Marcano & J. Cicero, #12290 (IIBZ); 1 ♂, Prov.
San Cristóbal, Campus Loyola, 28-xi-1973, E. Marcano #14910 (IIBZ); 1 ♂, 1 ♀, Prov. Peravia, Baní, Fundación
Sabana Buey, 1-vi-1972, E. Marcano & J. Cicero, #12289 (IIBZ); 1 ♂, Prov. Pedernales, entre Oviedo y
Pedernales, 24-iii-1967, E. Marcano, #4439 (IIBZ); 1 ♀, Prov. La Vega, Pontón, 26-iii-1970, E. Marcano, #6403
(IIBZ); 1 ♀, Prov. San Cristóbal, 17-iii-1970, E. Marcano & A. Abud, #6757, on batata (IIBZ); 1 ♀, Prov. San
Cristóbal, Campus Loyola, 10-xi-1970, E. Marcano, #8260 (IIBZ); 1 ♀, Prov. Peravia, Baní, Galeón, prox. Las
Carreras, xii-1970, E. Marcano, #9350 (IIBZ); 1 ♀, Prov. Pedernales, Villa del Mar, 7-viii-1971, E. Marcano & J.
Cicero, #10272 (IIBZ); 1 ♀, Prov. Puerto Plata, La Cumbre, 3-viii-1974, E. Marcano #17145 (IIBZ); 1 ♀, Prov.
San Cristóbal, Campus Loyola, 16-xii-1971, E. Marcano, #10505 (IIBZ); 1 ♀, Prov. Azua, Peralta, Puerta Vieja,
10-xi-1971, E. Marcano & J. Cicero, #9914 (IIBZ).
Catorhintha selector Stål

(Fig. 22)
Catorhinta selector Stål, 1859: 471.
Discussion. This species can be distinguished by having the antenniferous tubercle unarmed; antennal segments I
and III black; legs pale yellow and scattered with few black dots; thorax pale yellow with one black discoidal spot
on each segment; humeral angles obtuse; and dorsal abdominal segments black.
General distribution. Cited from Colombia, Costa Rica, Cuba, El Salvador, Guatemala, Haiti, Honduras,
Mexico, Nicaragua, the United States and Venezuela. HISPANIOLA: HAITI: La Serre (Brailovsky & Garcia 1987,
Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 4 ♂♂, Angostura, Peravia
Prov., 18°20.561’N 70°25.389’W, 27-x-2012, 215 m, D. Perez, B. Hierro, R. Bastardo (USNM). 4 ♀♀, Angostura,
Peravia Prov., 18°20.561’N 70°25.389’W, 27-x-2012, 215 m, D. Perez, B. Hierro, R. Bastardo (USNM). 4 ♀♀, El
Matadero, Peravia Prov., ~600 m, 30-viii-2014, D. E. Perez (USNM). 3 ♀♀, Azua Prov., on road to Las Yayitas,
175 m, 18°28.532’N 070°40.572’W, 26-VII-2017, T. Henry, M. Guidoti & D. Perez (USNM). 3 ♂♂, 4 ♀♀, Prov.
Azua, El Número, 22-vii-1979, Domínguez, Marcano, Reynoso (MNHNSD). 2 ♂♂, 1 ♀, 3-xii-78, Domínguez
(MNHNSD). 1 ♀, 12-x-1980, Domínguez (MNHNSD). 1 ♀, Prov. Azua, Tábara Abajo, 10-xi-1979, Marcano,
Cicero, Abud (MNHNSD). 1 ♀, Prov. Peravia, Cruce de Ocoa, 16-vi-1979, Marcano (MNHNSD). 1 ♂, San Juan,
15 km E San Juan, 8-viii-1989, C. W. O’Brien (UNAM); 2 ♂♂, Prov. Barahona, Laguna Caballero, 25-i-1976, E.
Marcano & J. Cicero, #19463 (IIBZ); 2 ♂♂, Prov. Barahona, Laguna Caballero, 25-i-1976, E. Marcano & J.
Cicero, #19462. on Plumbago (IIBZ); 1 ♂, Prov. Monte Cristi, Hatillo Palma, 8-ix-1965, E. Marcano, #2272
(IIBZ); 1 ♂, Prov. Peravia, Baní, Honduras, Sabana Roble, 5-vi-1971, E. Marcano & J. Cicero, #10166 (IIBZ); 2
♂♂, Prov. Peravia, Baní, Honduras, Sabana Roble, 5-vi-1971, E. Marcano & J. Cicero, #10174 (IIBZ); 1 ♀, Prov.
Peravia, Baní, Fundación Sabana Buey, 1-vi-1972, E. Marcano & J. Cicero, #12290 (IIBZ); 2 ♀♀. Prov. Peravia,
Baní, Galeón near Las Carreras, xii-1970, E. Marcano, #9350 (IIBZ). HAITI:1 ♂, 1 ♀, Santiago de los Santos, 6-
viii-1935, P. Thunb. (ZIZM).
Sethenira Spinola 1837
Diagnosis. Head not declivent; antenniferous tubercle unarmed; tylus compressed, raised above jugum: antennal
segment I thickened, curved, segments II–III dorso-ventrally flattened, IV shorter and rounded; anterolateral
margins of pronotum tuberculate; humeral angles broadly produced, truncated, and raised; triangular process
present; femora unarmed; abdomen moderately dilated on each side.
Distribution. A small genus with five species, represented throughout the Neotropics (Brailovsky 1988,
Packauskas 2010). In this contribution the genus is cited for the first time from Hispaniola.
Sethenira ferruginea Stål

(Fig. 23)
Sethenira ferruginea Stål, 1870: 182.
Discussion. Moderately large 14–20 mm, uniformly reddish brown to dark castaneous orange; rostrum reaching to
or beyond mesosternum; anterolateral margins of pronotum deeply concave; femora not swollen; and tibiae yellow
with irregular pale brown to black markings.
General distribution. Originally described from Cuba, and later cited from Bahamas, Belize, Mexico, and the
United States (Brailovsky 1988, Packauskas 2010). This species is cited for the first time from Hispaniola.
Material examined. New records. HISPANIOLA. DOMINICAN REPUBLIC: 3 ♂♂ 1 ♀, Pedernales,
Cabo Rojo, 10 m, 17°55’N 71°39’W, 1-x-91, R. Davidson, C. Young (UNAM); 1 ♀, Pedernales, km 22 carr. Cabo
Rojo-Aceitillar, Casa del Sr. Michell, (18°6’6”N 71°38’65”W), 513 m, 23-vii-2006. S. Vélez, B. Geller, J.
Henríquez, A. Marmolejo, R. Bastardo. Bosque latifoliado, lámpara vapor de mercurio (IIBZ).
Zicca Amyot and Serville 1843
Diagnosis. Medium sized bugs usually under 11 mm; ovobate or somewhat elongate and dull colored. Tylus
projected forward of bases of antennae, not compressed. Eyes large globular. Frontal region of cephalic disk with a
very deep, short longitudinal furrow. Pronotum strongly declivent, anterolateral margins tuberculate or spinose.
Humeral angles spinose. Hind femora clavate, swollen, and bearing ventral spines on distal half. Costal margin of
corium and upper margin of connexivum finely serrate. Abdomen not swollen. Body not dorsoventrally flattened.
Distribution. Primarily Neotropical, containing 19 species and one subspecies, most of which confined to
South America. In Hispaniola only one species (Z. taeniola [Dallas]) was previously cited. In this contribution two
species are added: Z. rubricator rubricator (Fabricius) and a new species here described (Brailovsky & Cadena
1992, Packauskas 2010).
Key to Hispaniola Species of Zicca
1. Connexivum pale orange with only posterior angle black; meso-, and metapleura without black discoidal spot; antennal seg-
ment IV yellow to yellowish orange with basal third black . . . . . . . . . . . . . . . Z. rubricator rubricator Brailovsky and Cadena
- Connexivum bicolorous, anterior half yellow, posterior half black; meso-, and metapleura with large black discoidal spot;
antennal segment IV entirely yellow to pale reddish orange . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
2. Fore, middle and hind femora entirely yellow; endocorium without white discoidal spot; posterior third of corium yellowish
white with apical angle dark brown; anterolateral margins of pronotum with small dark castaneous brown tubercles. . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Z. gloriosa Brailovsky and Perez-Gelabert sp. nov.
- Fore, middle and hind femora yellow with numerous black dots; middle third of endocorium with small white discoidal spot;
anterolateral margins of pronotum with a series of large white spines . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Z. taeniola (Dallas)
Zicca gloriosa Brailovsky & Perez-Gelabert sp. nov.

(Fig. 24)
Description. Holotype male. Dorsal coloration. Head dark orange, area around ocellar tubercle black; eyes
reddish brown; antennal segments I–III dark yellow, IV dark castaneous with apex dark yellow; pronotal disk pale
yellowish orange with the area around calli and two irregular longitudinal stripes at each side of midline black;
humeri, humeral angles and posterior margin dark castaneous brown; scutellum dark orange, punctures and two
basal discoidal spots lateral to midline black; apex dark yellow; clavus dark orange, punctures reddish; basal half of
corium dark yellow with punctures dark reddish; posterior half of corium pale yellowish white, apical angle dark
brown, punctures dark reddish and endocorium with dark castaneous irregular spot; hemelytral membrane dark
brown; connexival segments III–VI dark orange, with anterior third pale yellow, VII pale orange; abdominal
segments pale orange. Ventral coloration. Head pale orange; rostral segments dark yellow (apex of IV brown);
thorax pale orange; propleura with short black stripe near the anterolateral margins; meso- and metapleura each
with one black discoidal spot lateral to midline; anterior lobe of metathoracic peritreme dark orange, posterior lobe
pale orange; coxae orange with posterior margin dark yellow; trochanters, femora, tibiae and tarsi dark yellow;
abdominal sterna and genital capsule orange; sternites III–VI with a row of black dots on each side. Structure.
Head wider than long; antenniferous tubercle unarmed; tylus projecting well forward of antennae, not compressed;
eyes large, globular; ocellar tubercle raised; rostrum reaching anterior margin of metasternum; pronotum strongly
declivent; collar present; anterior border smooth; anterolateral borders with few tiny tubercles; humeral angles
projecting on a large spine directed laterally; upper third of posterolateral borders with few tiny tubercles, inner
third smooth; posterior border smooth; scutellum longer than wide, apically subacute; scutellar disk convex, gently
raised; fore and middle femora unarmed; hind femur relatively slender, armed with ventral spines at distal half;
macropterous; hemelytral membrane reaching the apex of last abdominal segment. Pronotum, scutellum, clavus,
corium, pro-, meso-, and metapleura densely and abruptly punctate; head, pro-, meso- and metasternum, and
abdominal sterna impunctate or with few scattered punctures. Body surface mostly glabrous.
Female. Unknown.
Measurements. Male. Body length 11.08; head length 1.45; head width across eyes 1.80; interocular distance
0.95; preocular distance 0.81; interocellar distance 0.39; antennal segments: I 2.00, II 2.40, III 1.95, IV 2.25;
pronotum: total length 2.35, maximum width across humeral angles 4.45; scutellum: length 1.44, width 1.25.
Type material. HOLOTYPE ♂: HISPANIOLA: DOMINICAN REPUBLIC: Santo Domingo Prov., Sierra
Prieta, lower hill, 140 m, 18°39.018’N–69°58.356’W, 23-viii-2016, D. Perez, S. Navarro, J. Borbón (UV light)
(USNM).
Etymology. From the Latin, “gloriosus”, referring to the attractive appearance of this species.
Discussion. Resembling Z. taeniola (Dallas) in having the connexivum bicolorous, and meso-, and metapleura
with black discoidal spot. In Z. gloriosa Brailovsky & Perez sp. nov., fore, middle and hind femora are entirely
yellow (versus yellow with black dots); endocorium without white discoidal spot (versus small white spot in the
middle of the corium); posterior third of corium yellowish white with apical angles brown (versus without
yellowish white area); and anterolateral margins of pronotum with small dark castaneous brown tubercles (versus
large white spines).
Zicca rubricator rubricator (Fabricius)

(Fig. 25)
Coreus rubricator Fabricius, 1803: 197.
Discussion. In Zicca rubricator rubricator (Fabricius 1803) the connexivum is nearly uniformly pale orange,
leaving only the posterior angle black; meso and metapleura without black discoidal spot; antennal segment IV
yellow to yellowish orange with basal third black; humeral angles produced into long acute spine directed outward;
and abdominal sterna without a row of black discoidal spots on each side. This subspecies is widely distributed
from Colombia to Brazil and across the transandine region from Ecuador to Bolivia (Brailovsky & Cadena, 1992).
The other known subspecies Zicca rubricator mayorgaea Brailovsky & Cadena (1992) occurs from southeastern
Mexico to Panama.
General distribution. Described from “America Meridionali” and later cited from Bolivia, Brazil, Colombia,
Costa Rica, Ecuador, El Salvador, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Surinam, and Venezuela
(Fabricius 1803, Brailovsky & Cadena 1992, Packauskas 2010). This species is cited for the first time from
Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♂♂, 3 ♀♀, Nuevos
Tablones, PN Armando Bermúdez, La Vega Prov., 1290 m, 19°02.946’N 70°53.298’W, 3-xi-2012, D. Perez, C.
Ramirez (uv light) (USNM); 1 ♂, Prov. San Cristóbal, La Colonia, Manomatuey, 18 29.512’N 70 15.082’W, 542
m, 6-ix-2008, R. Bastardo, D. Perez (IIBZ).
Zicca taeniola (Dallas)

(Fig. 26)
Clavigralla taeniola Dallas, 1852: 514.
Discussion. This is the most common species of Zicca in Hispaniola, characterized by having the endocorium with
small white spot; anterolateral margins of pronotum with a series of large white teeth; anterior lobe of pronotal disk
with two small white teeth or tubercles; mesopleura and metapleura each with one black discoidal spot; connexival
segments bicolorous, anterior half yellow, and posterior half black; fore, middle and hind femora with numerous
black dots; coxae with two large black spots on outer sides; antennal segment IV pale reddish orange (eventually
with apex pale yellow); and humeral angles with large and acute produced spine directed outward and slightly
forward.
General distribution. Described from Venezuela without formal localities and later cited from Belize,
Colombia, Costa Rica, Cuba, the Dominican Republic, Ecuador, El Salvador, Guatemala, Grenada, Honduras,
Jamaica, Mexico, Nicaragua, Panama, Puerto Rico, St. Vincent, Trinidad, the United States, and Venezuela.
HISPANIOLA: DOMINICAN REPUBLIC: San Cristóbal (Dallas 1852, Brailovsky & Cadena 1992, Packauskas
2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♂, Engombe, Santo
Domingo Oeste, 18°27.038’N 69°59.800’W, 20-viii-2014, D. Perez, S. Navarro (USNM). 1 ♂, Cerro Gordo, Baní,
Peravia Prov., 44 m, 18°16.067’N 70°20.603’W, 18-x-2012, D. Perez, R. Bastardo, A. Sanchez (USNM). 1 ♂, 19
kms NE Ocoa on road to Rancho Arriba, Ocoa Prov., 720 m, 18°39.003’N 70°27.849’W, 3-iv-2004, D. Perez, B.
Hierro, RD-209 (USNM). 2 ♀♀, HRD-026 Engombe, nr. UASD experimental fields, Santo Domingo Oeste, 32 m,
18°27.397’N 70°00.150’W, 05-viii-2015, D. Perez, S. Navarro, J. Sánchez B. (day+UV) (USNM). 1 ♀, Palacio de
Engombe, Santo Domingo, 18°27.047’N 69°59.802’W, 05-viii-2011, D. Perez, S. Navarro (USNM). 1 ♀,
Engombe, Santo Domingo Oeste, 18°27.038’N 69°59.800’W, 20-viii-2014, D. Perez, S. Navarro (USNM). 1 ♂, 1
♀, La Vega, nr. mouth Arroyo Los Dajaos, 5 km E. Manabao, 740 m, 19-04N 70-45W, 9-x-91, J. Rawlins, R.
Davidson, C. Young, S. Thompson (UNAM). 2 ♂♂, San Cristóbal, 6-vii-76, H. Ludovino (UNAM). 4 ♂♂, 3 ♀♀,
San Cristóbal, 24-i-76, H. L. Domínguez (UNAM). 1 ♂, San Cristóbal, 16-viii-76, H. Ludovino (UNAM). 1 ♂, 1
♀, Prov. El Seibo, Pedro Sánchez, 11-vi-76, E. E. Grissell (FSCA). Biological note: Sweeping grass. 1 ♀, 25 km N
Cabo Rojo, 700 m, 12-vii-1996, M. C. Thomas (FSCA). 1 ♂, La Vega Prov., 10 km W Jima, 24-v-1992, R.
Turnbow (UGAG). 1 ♂, La Vega Prov. Constanza, 17-vii-1996, R. Turnbow (UGAG). 9 ♂♂, 6 ♀♀, Prov. Peravia,
San José de Ocoa, Las Avispas, 15.viii.79, Domínguez y Marcano (MNHNSD).7 ♂♂, 16 ♀♀, Prov. El Seibo,
Sabana de la Mar, El Valle, Rio Yabón, 13-i-1980, E. Marcano (MNHNSD). 1 ♂, 1 ♀, Prov. La Altagracia, Higuey,
Veron, 10-viii-1979, E. Marcano, Mota, Reynoso (MNHNSD). 1♀, Prov. La Altagracia, Higuey, Punta Cana, 3-iv-
1980, Domínguez (MNHNSD). 1 ♂, 1 ♀, Prov. Puerto Plata, La Cumbre, 3-xi-79, Marcano y Mtz. (MNHNSD). 4
♂♂, 28-xii-1978, Domínguez, Silfa (MNHNSD). 1 ♀, Prov. Duarte, S. F. Macorís, La Guama, 26-iv-80, Reynoso
(MNHNSD). 6 ♂♂ 5 ♀♀, Prov. San Juan, L. M. Farfán, Las Carreras, 27-xii-1982, Marcano (MNHNSD). 1 ♀,
Prov. La Altagracia, Higuey, Juanillo, 21.iii.1981, Abud y Marcano (MNHNSD). 4 ♂♂, 4 ♀♀, Prov. Salcedo, Las
Cuevas, 18-vii-1979, Reynoso (MNHNSD). 1 ♀, Prov. El Seibo, Miches, Laguna El Limón, 29-iv-1980, Aquino y
Reynoso (MNHNSD). 1 ♀, Prov. Stgo. Rodríguez, Sabaneta, Gurabo, 5-vii-1980, Abud y Mota (MNHNSD). 1 ♀,
Prov. Santiago, Tamboril, Licey al Medio, 29-ix-1979, Marcano (MNHNSD). 2 ♀♀, San Juan, Sierra de Neiba,
Salcedo, caseta, 7.8 km SSW El Cercado, 18°39’56”N 71°51’57”W, 1659 m, 19-vi-2003, J. Rawlins, C. Young, R.
Davidson, C. Nuñez, P. Acevedo, M. de la Cruz (CMNH). Biological note: Pastures, pine Woodland, hand
collected. 1 ♀, Monseñor Nouel, Bonao, Presa Rio Blanco, 13-v-2001, C. A. Nuñez (MNHNSD). Note: Noct. 1 ♂,
Pedernales, 1 km S Los Arroyos, 1125 m, 18°14’N 71°45’W, 18-x-1991, R. Davidson, C. Young, S. Thompson, J.
Rawlins (CMNH). Biological note: Second growth forest. 6 ♂♂, 4 ♀♀, Santiago Prov., Babosico, 500 m,
19°20.983’N 70°37.447’W, 12-vii-2015, D. Perez, S. Navarro (day) (USNM); 3 ♂♂, 2 ♀♀, Prov. San Cristóbal,
Campus Loyola, 29-i-1971, E. Marcano, #8684 (IIBZ); 2 ♂♂, Prov. La Romana, Batey Diego, 18-vi-1970, E.
Marcano & R. Hansen, #7660 (IIBZ); 1 ♂, Prov. San Cristóbal, 19-ii-1975 E. Marcano, #17592 (IIBZ); 1 ♂, Prov.
Valverde, Esperanza, E. Marcano, #3163 (IIBZ); 2 ♂♂, 2 ♀♀, Prov. Pedernales, Agua Negra, 22-iii-1967, E.
Marcano, #3890 (IIBZ); 4 ♂♂, Prov. San Cristóbal, El Cacao, 450m, 9-i-1966, E. Marcano, #2983 (IIBZ); 1 ♂,
Prov. Santo Domingo, Campus UASD, 11-ix-1970, E. Marcano, #7763 (IIBZ); 1 ♂, Prov. La Vega, Jarabacoa, El
Pedregal, 29-vii-1967, E. Marcano, #4267 (IIBZ); 1 ♀, Prov. La Romana, Batey Diego, 18-vi-1970, E. Marcano &
R. Hansen, #7659 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 28-v-1967, E. Marcano, #4120 (IIBZ); 1 ♀,
Prov. La Romana, Batey Diego, 18-vi-1970, E. Marcano & R. Hansen, #7662 (IIBZ); 1 ♀, Prov. La Romana, Batey
Diego, 18-vi-1970, E. Marcano & R. Hansen, #7661 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 5-xii-1970,
E. Marcano, #8280 (IIBZ); 2 ♂♂, 1 ♀, Prov. Valverde, Esperanza, Boca de Mao, 7-iv-1971, E. Marcano & J.
Cicero, #8818 (IIBZ); 2 ♂♂, Prov. San Cristóbal, 17-iii-1970, E. Marcano, #6708 (IIBZ); 1 ♂, 2 ♀♀, Prov.
Valverde, Esperanza, Boca de Mao, 7-iv-1971, E. Marcano & J. Cicero, #8839 (IIBZ); 2 ♂♂, 1 ♀, Prov. Valverde,
Esperanza, Boca de Mao, 7-iv-1971, E. Marcano & J. Cicero, #8822 (IIBZ); 1 ♂, 2 ♀♀, Prov. Elías Piña, Carr.
Internacional, Pirámide 99, 450 m, 9-iii-1974, E. Marcano #15810 (IIBZ); 1 ♂, 1 ♀, Prov. Elías Piña, Carr.
Internacional, Pirámide 99, 450 m, 9-iii-1974, E. Marcano #15809 (IIBZ); 2 ♂♂, 1 ♀, Prov. San Pedro de Macorís,
Ramón Santana, Paso al Medio, 1977 E. Marcano, #21379 (IIBZ); 1 ♂, Prov. San Cristóbal, 30-iv-1976, E.
Macano, #19961 (IIBZ); 1 ♂, Prov. San José de Ocoa, Mahoma, 5-ii-1978 E. Marcano, #21943 (IIBZ); 1 ♂, Prov.
San Cristóbal, Campus Loyola, 16-xii-1971, E. Marcano, #10446 (IIBZ); 1 ♂, Prov. La Vega, Constanza, La
Descubierta, 8-ii-1971, E. Marcano & J. Cicero, #9428 (IIBZ); 1 ♂, 4 ♀♀, Prov. San Cristóbal, 30-iv-1976, E.
Macano, #19960 (IIBZ); 1 ♂, Prov. San Pedro de Macorís, Ramón Santana, Paso al Medio, 1977 E. Marcano,
#21388 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 4-ii-1972, E. Marcano, #10921 (IIBZ); 1 ♂, Prov. El
Seibo, Miches, Río Guarón, 2-ii-1975, E. Marcano & B. Monción, #17807 (IIBZ); 1 ♂, 1 ♀, Prov. La Vega,
Pontón, 26-iii-1970, E. Marcano, #6425 (IIBZ); 1 ♂, Prov. San José de Ocoa, Mahoma, 5-ii-1978 E. Marcano,
#21942 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 8-vi-1972, E. Marcano, #12351 (IIBZ); 1 ♂, Prov. San
Cristóbal, 17-iii-1970, E. Marcano, #6708 (IIBZ): 1 ♂, 1 ♀, Río Cumayasa, junto al puente La Roman-S.P. de
Macorís, 26-i-1973, E. Marcano #13112 (IIBZ); 2 ♀♀, Prov. Elías Piña, Carr. Internacional, Pirámide 99, 450 m, 9-
iii-1974, E. Marcano #15808 (IIBZ); 2 ♀♀, Prov. San José de Ocoa, Nizao, 16-v-1976 E. Marcano, #20262 (IIBZ);
2 ♀♀, Prov. San Cristóbal, Campus Loyola, 17-iii-1970, E. Marcano, #6633 (IIBZ); 1 ♀, Prov. San José de Ocoa,
Bejucal, los Palos Mojados, 11-iv-1976, E. Marcano & J. Cicero, #19887 (IIBZ); 4 ♀♀, Prov. El Seibo, Miches,
Río Guarón, 2-ii-1975, E. Marcano & B. Monción, #17829 (IIBZ); 1 ♀, Prov. San Cristóbal, 25-x-1972, E.
Marcano. #12786 (IIBZ); 1 ♀, Prov. Valverde, Mao, 4-vi-1973 E. Marcano #13602 (IIBZ); 2 ♀♀, Prov. San José
de Ocoa, El Pinal, 10-viii-1974 E. Marcano #17060 (IIBZ); 1 ♀, Prov. San José de Ocoa, Mahoma, 5-ii-1978 E.
Marcano, #21943 (IIBZ); 1 ♀, Prov. Elías Piña, Hondo Valle, Rancho de la Guardia, nac. Río Caña, 7-viii-1970, E.
Marcano & B. Patterson, #7838 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 16-xii-1971, E. Marcano,
#10444 (IIBZ); 1 ♂, Prov. La Romana, Chavón Arriba, Río Chavón, 31-iii-1973, E. Marcano #13274 (IIBZ); 1 ♂,
Prov. San Cristóbal, Campus Loyola, 10-xi-1970, E. Marcano, #8252 (IIBZ); 1 ♂, Prov. La Romana, Batey Diego,
18-vi-1970, E. Marcano & R. Hansen, #7657 (IIBZ); 1 ♂, Prov. Elías Piña, Hondo Valle, Rancho de la Guardia,
nac. Río Caña, 7-viii-1970, E. Marcano & B. Patterson, #7839; 1 ♂, Prov. Santo Domingo, Guerra, El Toro, 23-i-
1971, E. Marcano & H. Alain, #9173 (IIBZ); 1 ♂, Prov. Valverde, Barranca Bajón, Charco Feo, 19-iii-1977, E.
Marcano, #21279 (IIBZ); 1 ♂, Prov. San Cristóbal, Río Nigua, junto Iglesia San Gerónimo, 3-vi-1972, E. Marcano
& A. Abud, #12399 (IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 4-ii-1972, E. Marcano, #10922 (IIBZ); 1 ♂,
Prov. La Vega, Constanza, Cercadillo, junto arroyo Cabulla, 17-iv-1971, E. Marcano & S. Inchaustegui, #8925
(IIBZ); 1 ♂, Prov. San Cristóbal, Campus Loyola, 23-i-1974, E. Marcano #15630 (IIBZ); 1 ♂, Prov. Santiago,
Semana Santa, iii-1970, E. Marcano & L.Guzmán, #6834 (IIBZ); 1 ♂, Prov. Barahona, Polo, La Lanza, 25-iii-
1977, E. Marcano, #3997 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 4-ii-1972, E. Marcano, #10925 (IIBZ);
1 ♀, Prov. San Cristóbal, Campus Loyola, 16-xii-1971, E. Marcano, #10399 (IIBZ); 1 ♀, Prov. San Cristóbal,
Campus Loyola, 24-vii-1974, E. Marcano #16939 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 16-xii-1971,
E. Marcano, #10470 (IIBZ); 1 ♀, Prov. Santo Domingo, Engombe, 18-i-1970, E. Marcano & A. Abud, #5603
(IIBZ); 1 ♀, Prov. San Cristóbal, Campus Loyola, 4-ii-1972, E. Marcano, #10920 (IIBZ); 1 ♀, Prov. San Cristóbal,
17-iii-1970, E. Marcano & A. Abud, #6755. en batata (IIBZ); 1 ♀, San Cristóbal, San Juan, Mijo, Río Mijo, cruce
de la carret. 27-iii-1970, E. Marcano & G. Maldonado, #6454 (IIBZ); 1 ♀, Prov. La Romana, Chavón Arriba, Río
Chavón, 31-iii-1973 E. Marcano #13283 (IIBZ); 1 ♀, Prov. San Pedro de Macorís, Ramón Santana, Paso al Medio,
1977, E. Marcano, #21397 (IIBZ); 1 ♀, Prov. La Vega, Constanza, Arroyo del Toro, Quebrada Honda, 19-vii-1965,
E. Marcano, #1683 (IIBZ); 1 ♀, Prov. La Romana, Chavón Arriba, Río Chavón, 31-iii-1973, E. Marcano #13292
(IIBZ); 1 ♀, Prov. La Romana, Chavón Arriba, Río Chavón, 31-iii-1973, E. Marcano #13290 (IIBZ); 1 ♀, Prov. La
Romana, Batey Diego, 18-vi-1970, E. Marcano & R. Hansen, #7658 (IIBZ); 1 ♀, Prov. San Cristóbal, Campus
Loyola, 3-vi-1971, E. Marcano, #10346 (IIBZ). HAITI: 3 ♂♂, 2 ♀♀, Port. au Prince, vi, E. D. Ball, 1940,
(USNM).
Tribe Spartocerini
Key to Hispaniola Genera of Spartocerini
1. Anterolateral margins of pronotum with short to large nodular processes; antennal segment I shorter than 1.80 mm; antennal
segment IV shorter than 2.15 mm; pronotal disk with a transverse sinuated carina at the middle, sometimes hard to see…
Eubule Stål . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .E. spartocerana Brailovsky
- Anterolateral margins of pronotum smooth, without spines or nodular processes; antenal segment I longer than 2.40 mm;
antennal segment IV longer than 2.65 mm; pronotal disk without transverse carina . . . . . . . . . . . . . . . . . . . Spartocera Laporte
Eubule Stål 1867
Diagnosis. Medium size species. Head quadrate, conspicuously declivent. Antennal segments short, clothed with
thick golden setae. Buccula short, raised, reaching anterior margin of eyes. Pronotum declivent. Anterolateral
margins of pronotum with short to large nodular processes. Humeral angles slightly exposed. Pronotal disk with a
transverse, sinuated carina at the middle, sometimes hard to see. Forewing with numerous anastomosing veins in
the membrane. Legs unarmed.
Distribution. The genus includes 13 species distributed through the Greater and Lesser Antilles, and South
America (Brailovsky 1987, 1992, Packauskas 2010).
Eubule spartocerana Brailovsky

(Fig. 27)
Eubule spartocerana Brailovsky, 1992: 209–212.
Discussion. Body dorsally and ventrally including the legs densely clothed with erect and decumbent silvery and
gloden setae; rostrum reaching anterior border of mesosternum; rostral segment I reaching anterior margin of
prosternum; scutellum black with yellowish longitudinal stripe along midline; corium dark to pale castaneous
orange with apical margin and short irregular stripe between midlle third of endocorium and apical margin black to
dark brown; hemelytral membrane pale yellow with numerous anastamosing veins; pro-, meso-, and metapleura
pale castaneous orange with five shiny black elongate spots, two on propleura and mesopleura and one on
metapleural; and connexival margins IV–VIII black to dark brown with anterior border irregularly yellow.
General distribution. Only cited from Surinam and Venezuela (Brailovsky 1992, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 1 ♀, Independencia, Sierra
de Bahoruco, north slope, 13.5 km SE Puerto Escondido, 18°12’18”N 71°31’08”W, 1789 m, 24–25-xi-2004, J.
Rawlins, C. Young, C. Nuñez, V. Verdecia, W. A. Zanol (CMNH). 12 ♂♂ 16 ♀♀ and 87 nymphs, República
Dominicana, San Cristóbal, Nigua, Puerta Blanca, 5.ix.2018, colectores: Juana Peña, Cipriano Rosario, General
Segura (IIBZ).
Spartocera Laporte 1832
Diagnosis. Medium to large broad, ovoid species. Head quadrate, conspicuously declivent. Antennal segments
relatively short, and clothed with thick black setae. Pronotum declivent. Collar present. Anterolateral margins of
pronotum sinuate and nodulose. Humeral angles prominent, gently rounded, or truncated. Posterior border slightly
convex. Forewing with numerous anastomosing veins in the membrane. Femora unarmed. Metathoracic peritreme
bilobate. Abdomen markedly dilated. Connexivum raised above terga.
Distribution. Large genus with 18 species widely distributed from the United States throughout Mexico,
Central America, Greater and Lesser Antilles, and South America including Argentina and Chile (Brailovsky &
Sanchez 1983, Packauskas 2010). Two species occur in Hispaniola.
Key to Hispaniola Species of Spartocera
1. Femora black with basal joint and apical third shiny reddish orange; tibiae black with basal third shiny reddish orange; clavus
and corium usually yellow; scutellum yellow with basal angles and longitudinal medial stripe black; connexival segments III–
VII yellow with anterior margin black or only with upper border of anterior margin black; humeral angles rounded, exposed,
and directed upward . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S. fusca (Thunberg)
- Femora and tibia uniformly dark castaneous orange; clavus and corium dark castaneous orange; scutellum dark castaneous
orange with apex yellow; connexival segments III–VII dark castaneous orange with anterior and posterior third shiny reddish
orange or reddish orange with posterior border dark castaneous orange; humeral angles obtuse, scarsely exposed . . . . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .S. batatas (Fabricius)
Spartocera batatas (Fabricius)

(Fig. 28)
Lygaeus batatas Fabricius, 1798: 540.
Discussion. Recognized by pronotum black with anterolateral margins, humeral angles and short longitudinal
stripe at middle third of posterior lobe reddish to yellow; anterolateral margins of pronotum not expanded; humeral
angles generally bluntly rounded or obtuse, not exposed; clavus black to dark brown; corium black to dark brown
with tiny yellowish spot at middle third of endocorium; connexivum reddish with posterior border black.
General distribution. Cited from Brazil, Colombia, Cuba and Surinam (Packauskas 2010). This species is
cited for the first time from Hispaniola.
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♀♀, Santo Domingo
Norte, Mirador del Norte, x-2014, C. de Soto Molinari (USNM). 1 ♀, DR-2016-034, Pedernales Prov., Riosito,
18°00.024’N 71°11.031’W, 28-viii-2016, D. Perez, S. Medrano, M. Landestoy (USNM). 1 ♂, 1 ♀, Barahona,
Payaso, 13-vii-96, R. Turnbow (UGAG). 3 ♂♂, 2 ♀♀, Las Cuevas, Prov. Salcedo, Salcedo, 27-iv-1980, Reynoso
(MNHNSD). 1 ♂, Prov. San Juan, San Juan, 3-ii-1973, Domínguez (MNHNSD). 1♀, Distrito Nal., 4 km W Boca
Chica, 11-viii-1979, L. B. O’Brien (UNAM). 1 ♀, Santo Domingo, WI, 17-viii, Busck (USNM); 1 ♂, Prov.
Barahona, Enriquillo, xii-1969, E. Marcano, #5507 (IIBZ); 1 ♂, Prov. San Cristóbal, Sabana Grande de Boyá,
Majagual, 1-xii-1973, E. Marcano #14901 (IIBZ); 1 ♂, Santo Domingo, Los Minas, vii-1973, E. Marcano #14088;
1 ♀, Prov. San Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14897 (IIBZ); 1 ♀, Prov.
San Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14898 (IIBZ); 1 ♀, Prov. San
Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14899 (IIBZ).
Spartocera fusca (Thunberg)
(Fig. 29)
Cimex fuscus Thunberg, 1783: 44.
Discussion. This species is easily recognized by having the pronotum pale yellow with 2 to 4 black longitudinal
stripes. Additionally the anterolateral margins of pronotum are expanded; humeral angles distinctly raised,
truncate; clavus and corium pale yellow; legs black to dark brown with coxae, trochanter, basal and apical joint of
femora and basal joint of tibiae reddish; and connexival segments yellow with anterior third usually black.
General distribution. Widespread throughout the United States, Mexico, Central America, Greater and Lesser
Antilles and South America. Antigua, Argentina, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Dominica, the
Dominican Republic, French Guiana, Guadeloupe, Guatemala, Jamaica, Martinique, Mexico, Nicaragua, Panama,
Peru, Puerto Rico, St. Bartholomew, St. Martin, St. Vincent, the United States, Uruguay, and Venezuela.
HISPANIOLA: DOMINICAN REPUBLIC: without formal locality (Barber 1923a, Blöte, 1938, Brailovsky &
Sanchez 1983, Perez-Gelabert 2008, Packauskas 2010).
Material examined. New records. HISPANIOLA: DOMINICAN REPUBLIC: 2 ♂♂, Monseñor Nouel
Prov., rd. to Blanco hydroelectric, 18°52.940’N 70°30.346’W, 700 m, 13-viii-2016, D. Perez, R. Bastardo, A.
Sánchez (USNM). 1 ♀, Arroyo El Derrumbao, nr. confluencia w/Rio Yaque del Norte, La Vega Prov., 1006 m,
19°04.626’N 70°50.595’W, 2-xi-2012, D. Perez, C. Ramírez (USNM). 1 ♀, DR-2016-011, Monseñor Nouel Prov.,
Centro ecoturístico Rio Blanco, 18°52.635’N 70°30.391’W, 669 m, 13–14-viii-2016, D. Perez, R. Bastardo, A.
Sánchez (USNM). 1 ♀, DR-2016-007, San Cristóbal Prov. On dirt road going up, Rancho Viejo, 18°28.332’N
70°15.553’W, 950 m, 10-viii-2016, D. Perez, C. de Soto Molinari, D. Pou (USNM). 1 ♀, Masacó, Prov. Barahona,
743 m, 18°02.081’N’ 71°18.591’W, 16-viii-2011, D. Perez, S. Navarro, R. Gómez (day + UV light) (USNM). 1 ♀,
Peravia, Arroyo Cañas, 650 m, 7-viii-80, A. Norrbom (UNAM). 1 ♂, Prov. La Vega, La Vega, 24-iii-1972,
Domínguez (MNHNSD). 6 ♂♂, 8 ♀♀, Baoruco, Sierra de Neiba, Los Guineos, on upper Rio Colorado, 18°35’N
71°11’W, 630 m, 11–12-viii-1990, J. Rawlins, S. Thompson (CMNH). Biological notes: Feeding on Mikania
scandens. Mesic riparian woodland. 1 ♂, 10 km E Hondo Valle, La Estrelleta, 7-viii-1979, L. B. O’Brien (UNAM);
1 ♂, Prov. Santo Domingo, 24-x-1971, E. Marcano & G. Olivo, #10313 (IIBZ); 1 ♂, Prov. Pedernales, Banano, Río
Mulito, 17-viii-1973, E. Marcano #14097 (IIBZ); 1 ♂, Prov. San Cristóbal, Sabana Grande de Boyá, Majagual,
1.xii-1973, E. Marcano #14900 (IIBZ); 1 ♂, Prov. San Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E.
Marcano #14898 (IIBZ); 1 ♂, Prov. San Cristóbal, ciudad, 19-v-1976, E. Marcano, #20031 (IIBZ); 1 ♂, Prov.
Monte Plata, Bayaguana, Comatillo, Rincón Naranjo, 29-ii-1964, E. Marcano, #332 (IIBZ); 1 ♀, Prov. Santiago,
Tamboril, Licey al Medio, 15-iv-1973, E. Marcano #13173 (IIBZ); 1 ♀, Prov. San Cristóbal, 6-xii-1976, E.
Marcano, #20776 (IIBZ); 1 ♀, Prov. San Cristóbal, ciudad, 19-v-1976, E. Marcano, #20016 (IIBZ); 1 ♀, Prov.
Puerto Plata, La Cumbre, 6-viii-1974, E. Marcano #16762 (IIBZ); 1 ♀, Prov. San Cristóbal, 19-ii-1975, E.
Marcano #17595 (IIBZ); 1 ♀, Prov. Santiago, Licey al Medio, 20-x-1968, E. Marcano, #4816 (IIBZ); 1 ♀, Prov.
Santiago, Tamboril, Licey al Medio, 16-iv-1973, E. Marcano #13184 (IIBZ); 1 ♀, Prov. Elías Piña, Pirámide 204,
31-v-1975, E. Marcano & A. Abud, #18412 (IIBZ); 1 ♀, Prov. Santo Domingo, 17-v-1970, E. Marcano, #7216
(IIBZ); 1 ♀, Prov. Puerto Plata, La Cumbre, 6-viii-1974, E. Marcano #16759 (IIBZ); 1 ♀, Prov. La Vega,
Constanza, Yaquecillo, viii-1958, E. Marcano, #156 (IIBZ); 1 ♀, Prov. Santiago, Licey al Medio, 7-iv-1971, E.
Marcano, #9008 (IIBZ); 1 ♀, Prov. Santiago Rodríguez, 4-vii-1964, E. Marcano, #664 (IIBZ); 1 ♀, Prov. Santo
Domingo, Los Minas, 18-iv-1970, E. Marcano, #6909 (IIBZ); 1 ♀, Prov. Puerto Plata, La Cumbre, 6-viii-1974, E.
Marcano #16760 (IIBZ); 1 ♀, Prov. Santiago, Licey al Medio, 8-viii-1965, E. Marcano, #1895 (IIBZ); 1 ♀, Prov.
San Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14900 (IIBZ); 1 ♀, Prov. Santiago, San
José de las Matas, Mata Grande, Río Antosape, 4-iv-1970, E. Marcano, #6882 (IIBZ); 1 ♀, Prov. La Vega, Licey,
29-v-1972, E. Marcano, #12324 (IIBZ); 1 ♀, Prov. Pedernales, Agua Negra, 22-iii-1967, E. Marcano, #3874
(IIBZ); 1 ♀, Prov. San Cristóbal, La Cabirma de la Loma, 14-ii-1976, E. Marcano, #19605 (IIBZ); 1 ♀, Prov. San
Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14902 (IIBZ); 1 ♀, Prov. La Vega, Piedra
Blanca, El Cacao, Km 14, 23-x-1971, E. Marcano, #9926 (IIBZ); 1 ♀, Prov. Santiago, Licey al Medio, 20-x-1968,
E. Marcano, #4815 (IIBZ); 1 ♀, Prov. Samaná, Sánchez, Majagual, 12-iv-1965, E. Marcano, #1131 (IIBZ); 1 ♀,
Prov. Puerto Plata, Arroyo del Toro, Cañada Honda, 17-viii-1965, E. Marcano, #1928 (IIBZ); 1 ♀, Prov.
Independencia, Jimaní, Guzmán, 19-v-1973, E. Marcano #14761 (IIBZ); 1 ♀, Prov. Independencia, Jimaní,
Guzmán, 19-v-1973 E. Marcano #14762 (IIBZ); 1 ♀, Prov. Monte Plata, Arroyo Rancho, entre Río Verde y
Yamasá, 24-xi-1973, E. Marcano #14987, on Commelina virginica (IIBZ); 1 ♀, Prov. Barahona, Enriquillo, xii-
1969, E. Marcano, #5506 (IIBZ); 1 ♀, Prov. Santo Domingo, 10-x-1971, E. Marcano & B. García, #9673 (IIBZ); 1
♀, Prov. San Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14899 (IIBZ); 1 ♀, Prov. San
Cristóbal, Sabana Grande de Boyá, Majagual, 1-xii-1973, E. Marcano #14901 (IIBZ); 1 ♀, Engombe, Santo
Domingo, D. N. 4-iii-76, UASD, Algodón (IIBZ); 1 ♀, Prov. Sánchez Ramírez, Cotui, Fátima, 2087200-376223,
346 m, 25–26-vii-2008, R. Bastardo, D. Perez (IIBZ); 1 ♂, Santo Domingo Este, Parque Mirador Este, 15-iv-2017,
Y. Piña (IIBZ).
Acknowledgements
We are grateful to the following colleagues for access to or loan of specimens under their care: Mick Webb
(BMNH), Vincent Lee, Christopher C. Grinter and the late Norman Penny (CASC), Rosser Garrison (CDFA), John
Rawlins (CMNH), Philip P. Parrillo (FMNH), Susan Halbert and Julieta Brambila (FSCA), Ruth H. Bastardo
(IIBZ), Gabriel de los Santos (MNHNSD), Karen Wright and Joseph C. Schaffner (TAMU), Joe McHugh and Cecil
Smith (UGAG), Philip J. Clausen (UMSP), Thomas J. Henry (USNM), and H. Strumpel (ZIZM). Brígido Hierro
(Ministerio de Medio Ambiente, Santo Domingo), Arturo León Benítez and Francisco Paz (MNHNSD) and Judá
Isaí Martínez (IIBZ) assisted DEPG in Santo Domingo. Lucrecia H. Rodríguez (Systematic Entomology Lab,
USDA) provided technical expertise to improve the illustrations. Special thanks to Ernesto Barrera (UNAM) for
technical support and to Oscar Federico Francke Balle (UNAM) for providing a critical review of this manuscript.
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Check list of the known subfamilies, tribes, genera and species of Dominican Republic
Order Hemiptera Linnaeus, 1758

Suborder Heteroptera Latreille, 1810
Family Coreidae Leach, 1815
Subfamily Meropachyinae Stål, 1867
Tribe Merocorini Kormilev, 1954
Genus Merocoris Perty, 1930
Subgenus Corynocoris Mayr, 1866
M. (C.) distinctus distinctus Dallas, 1852
M. (C.) typhaeus (Fabricius, 1798)
Subfamily Coreinae Leach, 1815
Tribe Acanthocerini Bergroth, 1913
Genus Acanthocerus Palisot de Beauvois, 1818
A. crucifer Palisot de Beauvois, 1818
Tribe Anisoscelini Laporte, 1832
Genus Chondrocera Laporte, 1832
C. mirabilis Brailovsky & Barrera, 1994
Genus Leptoglossus Guerín-Méneville, 1831
L. balteatus (Linnaeus, 1771)
L. concolor (Walker, 1871)
L. confusus Alayo & Grillo, 1977
L. gonagra (Fabricius, 1775)
Genus Mamurius Stål, 1862
M. cubanus Barber & Bruner, 1947
Genus Phthia Stål, 1862
P. rubropicta (Westwood, 1842)
Genus Phthiacnemia Brailovsky, 2009
P. picta (Drury, 1770)
Genus Rhytidophthia Brailovsky, 2009
R. splendida (Valdes, 1910)
Tribe Chariesterini Stål, 1867
Genus Chariesterus Laporte, 1832
Ch. gracilicornis Stål, 1870
Tribe Hypselonotini Bergroth, 1913
Genus Althos Kirkaldy, 1904
A. obscurator (Fabricius, 1803)
Genus Anasa Amyot & Serville, 1843
A. acutangula Stål, 1870
A. bellator (Fabricius, 1787)
A. pisina Brailovsky, 2001
A. scorbutica (Fabricius, 1775)
A. tristis (De Geer, 1773)
Genus Catorhintha Stål, 1859
C. guttula (Fabricius, 1794)
C. selector Stål, 1859
Genus Sethenira Spinola, 1837
S. ferruginea Stål, 1870
Genus Zicca Amyot & Serville, 1843
Z. gloriosa sp. nov.
Z. rubricator rubricator (Fabricius, 1803)
Z. taeniola (Dallas, 1852)
Tribe Spartocerini Amyot & Serville, 1843
Genus Eubule Stål, 1867
E. spartocerana Brailovsky, 1992
Genus Spartocera Laporte, 1832
S. batatas (Fabricius, 1798)
S. fusca (Thunberg, 1783)
Check list of the known subfamiies, tribes, genera and species of Haiti
Order Hemiptera Linnaeus, 1758

Suborder Heteroptera Latreille, 1810
Family Coreidae Leach, 1815
Subfamily Coreinae Leach, 1815
Tribe Acanthocerini Bergroth, 1913
Genus Acanthocerus Palisot de Beauvois, 1818
A. crucifer Palisot de Beauvois, 1818
Tribe Anisoscelini Laporte, 1832
Genus Leptoglossus Guerín-Méneville, 1831
L. balteatus (Linnaeus, 1771)
L. gonagra (Fabricius, 1775)
Genus Rhytidophthia Brailovsky, 2009
R. splendida (Valdes, 1910)
Tribe Chariesterini Stål, 1867
Genus Chariesterus Laporte, 1832
Ch. gracilicornis Stål, 1870
Tribe Hypselonotini Bergroth, 1913
Genus Anasa Amyot & Serville, 1843
A. scorbutica (Fabricius, 1775)
Genus Catorhintha Stål, 1859
C. selector Stål, 1859
Genus Zicca Amyot & Serville, 1843
Z. taeniola (Dallas, 1852)
FIGURES 2–10. Habitus color pictures of the species of Coreidae recorded from Hispaniola. 2. Merocoris (Corynocoris)
distinctus, 3. Merocoris (Corynocoris) typhaeus, 4. Acanthocerus crucifer, 5. Chondrocera mirabilis, 6. Leptoglossus
balteatus, 7. Leptoglossus confusus, 8. Leptoglossus concolor, 9. Leptoglossus gonagra, 10. Mamurius cubanus. In all photos
scale represents approximately 5 mm.
FIGURES 11–20. Habitus color pictures of the species of Coreidae recorded from Hispaniola. 11. Phthia rubropicta, 12.
Phthiacnemia picta, 13. Rhytidophthia splendida, 14. Chariesterus gracilicornis, 15. Althos obscurator, 16. Anasa acutangula,
17. Anasa bellator, 18. Anasa pisina, 19. Anasa scorbutica, 20. Anasa tristis. In all photos scale represents approximately 5
mm.
FIGURES 21–29. Habitus color pictures of the species of Coreidae recorded from Hispaniola. 21. Catorhintha guttula, 22.
Catorhintha selector, 23. Sethenira ferruginea, 24. Zicca gloriosa sp. nov. 25. Zicca rubricator, 26. Zicca taeniola, 27. Eubule
spartocerana, 28. Spartocera batatas, 29. Spartocera fusca. In all photos scale represents approximately 5 mm.

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Zootaxa 4568 (2): 201–241 ISSN 1175-5326 (print edition)

A review of the Coreidae of Hispaniola (Hemiptera: Heteroptera),

HARRY BRAILOVSKY1 & DANIEL E. PEREZ-GELABERT2

Accepted by D. Rider: 8 Feb. 2019; published: 20 Mar. 2019 201

FIGURE 1. Map of the island of Hispaniola.

Material and methods

Key to Hispaniola subfamilies, tribes, genera and species of Coreidae* »»

*Modified from Packauskas key (1994)

Merocoris Perty 1830

Merocoris (Corynocoris) Mayr 1866

Key to Hispaniola species of Merocoris (Corynocoris)

Merocoris (C.) distinctus Dallas

Merocoris distinctus Dallas, 1852: 419.

Merocoris (C.) typhaeus (Fabricius)

Lygaeus typhaeus Fabricius, 1798: 537.

Acanthocerus Palisot de Beauvois 1818

Acanthocerus crucifer Palisot, 1818, 204.

Key to Hispaniola genera and species of Anisoscelini

1. Hind tibiae foliaceous, strongly expanded on both sides . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

Chondrocera Laporte 1832

Chondrocera mirabilis Brailovsky & Barrera, 1994: 54–57.

Leptoglossus Guérin-Méneville 1831

Key to Hispaniola species of Leptoglossus

Leptoglossus balteatus (Linnaeus)

Cimex balteatus Linnaeus, 1771: 534.

Anisoscelis concolor Walker, 1871: 128.

Leptoglossus confusus Alayo & Grillo

Leptoglossus confusus Alayo & Grillo, 1977: 97.

Leptoglossus gonagra (Fabricius)

Cimex gonagra Fabricius, 1775: 708.

Mamurius Stål 1862

Mamurius cubanus Barber & Bruner

Mamurius cubanus Barber & Bruner, 1947: 78–79.

Phthia Stål 1862

Phthia rubropicta (Westwood)

Leptoscelis rubropicta Westwood, 1842: 17.

Phthiacnemia Brailovsky 2009

Phthiacnemia picta (Drury)

Cimex picta Drury, 1770: 107.

Rhytidophthia Brailovsky 2009

Rhytidophthia splendida (Valdes)

Phthia splendida Valdes, 1910: 431.

Chariesterus De Laporte 1832

Chariesternus gracilicornis Stål

Chariesternus gracilicornis Stål, 1870: 178.

Key to Hispaniola Genera of Hypselonotini

1. Hind femora armed ventrally with two or more spines. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

Althos Kirkaldy 1904

Althos obscurator (Fabricius)

Coreus obscurator Fabricius, 1803: 200.

Anasa Amyot and Serville 1843

Key to Hispaniola Species of Anasa

1. Hind femora armed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . A. scorbutica (Fabricius) (partly)

Anasa acutangula Stål

Anasa acutangula Stål, 1870: 196–197.

Anasa bellator (Fabricius)

Cimex bellator Fabricius 1787: 286.

Anasa pisina Brailovsky, 2001, 43–45.

Anasa scorbutica (Fabricius)

Cimex scorbuticus Fabricius, 1775: 706.

Cimex tristis De Geer, 1773: 340.

Catorhintha Stål 1859

Key to Hispaniola Species of Catorhintha