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Molar (tooth)
TA A05.1.03.007
FMA 55638
Anatomical terminology
Human anatomy
In humans, the molar teeth have either
four or five cusps. Adult humans have 12
molars, in four groups of three at the back
of the mouth. The third, rearmost molar in
each group is called a wisdom tooth. It is
the last tooth to appear, breaking through
the front of the gum at about the age of
20, although this varies from individual to
individual. Race can also affect the age at
which this occurs, with statistical
variations between groups.[1] In some
cases, it may not even erupt at all.
Mammal evolution
In mammals, the crown of the molars and
premolars is folded into a wide range of
complex shapes. The basic elements of
the crown are the more or less conical
projections called cusps and the valleys
that separate them. The cusps contain
both dentine and enamel, whereas minor
projections on the crown, called
crenulations, are the result of different
enamel thickness. Cusps are occasionally
joined to form ridges and expanded to
form crests. Cingula are often incomplete
ridges that pass around the base of the
crown.[2]
Tribosphenic …
Generalized tribosphenic molar: The protocone is on
the lingual (tongue) side, while the anterior paracone
and posterior metacone are on the buccal (cheek)
side of the jaw).
Quadrate …
Pig tooth
Quadrate (also called quadritubercular or
euthemorphic) molars have an additional
fourth cusp on the lingual (tongue) side
called the hypocone, located posterior to
the protocone. Quadrate molars appeared
early in mammal evolution and are present
in many species, including hedgehogs,
raccoons, and many primates, including
humans.[7] There may be a fifth cusp.
Bunodont …
Upper and lower dentition of a chimpanzee
Hypsodont …
Hypsodont dentition is characterized by
high-crowned teeth and enamel that
extends far past the gum line, which
provides extra material for wear and tear.[9]
Some examples of animals with
hypsodont dentition are cattle and horses,
all animals that feed on gritty, fibrous
material. Hypsodont molars can continue
to grow throughout life, for example in
some species of Arvicolinae (herbivorous
rodents).[7]
Hypsodont molars lack both a crown and a
neck. The occlusal surface is rough and
mostly flat, adapted for crushing and
grinding plant material. The body is
covered with cementum both above and
below the gingival line, below which is a
layer of enamel covering the entire length
of the body. The cementum and the
enamel invaginate into the thick layer of
dentin.[10]
Brachydont …
The opposite condition to hypsodont is
called brachydont or brachyodont (from
brachys, "short"). It is a type of dentition
characterized by low-crowned teeth.
Human teeth are brachydont.[7]
Zalambdodont …
Dilambdodont …
Lophodont …
Lophodont molars of Elephas (left) and Loxodonta
(center), compared to the nonlophodont mastodon
(right)
Selenodont …
Secodont …
See also
Dental formula
Polyphyodont
Notes
1. Rozkovcová, E; Marková, M; Dolejsí, J
(1999). "Studies on agenesis of third
molars amongst populations of
different origin". Sbornik Lekarsky. 100
(2): 71–84. PMID 11220165 .
2. Zhao, Weiss & Stock 2000, Acquisition
of multi-cusped cheek teeth in
mammals, p. 154
3. Myers et al. 2013b
4. Stokstad 2001
5. Luo, Cifelli & Kielan-Jaworowska 2001
6. Luo, Ji & Yuan 2007
7. Myers et al. 2013a
8. Lawlor 1979, pp. 13–4
9. Flynn, Wyss & Charrier 2007
10. Kwan, Paul W.L. (2007). "Digestive
system I" (PDF). Tufts University.
Archived from the original (PDF) on
2012-09-13. Retrieved 2013-05-18.
References
Flynn, John J.; Wyss, André R.; Charrier,
Reynaldo (May 2007). "South America's
Missing Mammals" . Scientific American: 68–
75. OCLC 17500416 . Retrieved 2013-05-11.
Lawlor, T.E. (1979). "The Mammalian
Skeleton" (PDF). Handbook to the Orders and
Families of Living Mammals. Mad River Press.
ISBN 978-0-916422-16-5. OCLC 5763193 .
Archived from the original (PDF) on 2010-07-
01. Retrieved 2013-05-12.
Luo, Zhe-Xi; Cifelli, Richard L.; Kielan-
Jaworowska, Zofia (4 January 2001). "Dual
origin of tribosphenic mammals". Nature.
409 (6816): 53–7. doi:10.1038/35051023 .
PMID 11343108 .
Luo, Z.-X.; Ji, Q.; Yuan, C.-X. (November
2007). "Convergent dental adaptations in
pseudo-tribosphenic and tribosphenic
mammals". Nature. 450 (7166): 93–97.
doi:10.1038/nature06221 . PMID 17972884 .
Myers, P.; Espinosa, R.; Parr, C. S.; Jones, T.;
Hammond, G. S.; Dewey, T. A. (2013a). "The
Basic Structure of Cheek Teeth" . Animal
Diversity Web, University of Michigan.
Retrieved 2013-05-12.
Myers, P.; Espinosa, R.; Parr, C. S.; Jones, T.;
Hammond, G. S.; Dewey, T. A. (2013b). "The
Diversity of Cheek Teeth" . Animal Diversity
Web, University of Michigan. Archived from
the original on 2013-04-05. Retrieved
2013-05-12.
Stokstad, E. (January 2001). "Tooth Theory
Revises History of Mammals" . Science. 291
(5501): 26.
doi:10.1126/science.10.1126/science.291.5
501.26 . PMID 11191993 .
Zhao, Z.; Weiss, K. M.; Stock, D. W. (2000).
"Development and evolution of dentition
patterns and their genetic basis". In Teaford,
Mark F; Smith, Moya Meredith; Ferguson,
Mark WJ (eds.). Development, Function and
Evolution of Teeth. Cambridge University
Press. pp. 152–72. ISBN 978-0-511-06568-2.
External links
Overview of molar morphology and
terminology - Paleos.com
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