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WHEY PROCESSING

Contents
Utilization and Products
Demineralization

Utilization and Products


P Jelen, University of Alberta, Edmonton, AB, Canada
ª 2011 Elsevier Ltd. All rights reserved.

Introduction The main components of both sweet and acid wheys,


after water (which constitutes approximately 93% of
Whey, the greenish translucent liquid obtained from milk the whey on an ‘as is’ basis), are lactose (approximately
after precipitation of casein, has been viewed until recently 70–72% of the total solids), whey proteins (approximately
as one of the major disposal problems of the dairy industry. 8–10%), and minerals (approximately 12–15%). Table 1
The biological oxygen demand (BOD) of whey is very high gives a more detailed breakdown of these components of
(40 000 mg kg 1 or more), constituting a major ecological the two basic whey types. The main differences between
burden if disposed off as a waste material. Thus, the disposal the two whey types are in the mineral content, the acidity,
practices of the past, including drainage into waste treatment and the composition of the whey protein fraction (WPF).
facilities or spraying onto fields, are currently seldom prac- Although these differences are relatively minor on an ‘as is’
ticed. Use of whey as cattle or pig feed is still one of the basis, they can have a profound effect on the technological
significant alternatives to utilization in the human food as well as nutritional properties of the wheys and must be
chain, now being predominantly favored due to the eco- taken into consideration in applications of the various
nomic opportunities provided by some of the milk nutrients whey processing technologies now available to whey pro-
contained in the whey. cessors. The acid coagulation approach (using conversion
of some of the lactose in milk to lactic acid by lactic acid
bacteria and/or addition of acidulants such as glucono--
Whey Types and Composition lactone or various acids such as sulfuric, phosphoric,
hydrochloric, citric, or lactic acid) results in substantially
There are several types of whey, depending mainly on the increased acidity (final pH approximately 4.5) necessary
processing sequence resulting in casein removal from for casein precipitation. At this low pH, the colloidal
fluid milk. The type of whey most often encountered calcium contained in the casein micelles in normal milk
originates from the manufacture of cheese or certain is solubilized and partitioned into the whey. On the other
industrial casein products, where the processing is based hand, rennet clotting produces a fragment of the -casein
on coagulating the casein by rennet, an industrial molecule, termed glycomacropeptide (GMP), which ends
casein-clotting preparation containing chymosin or up in the whey. Thus, the GMP constitutes approximately
other casein-coagulating enzymes. Since the rennet- 20% of the WPF of sweet, rennet-based wheys but is not
induced coagulation of casein and the subsequent whey found in acid wheys unless use of rennet was included in
drainage occur at a pH value of approximately 6.5–6.0, the fresh cheese manufacturing process (as sometimes
this type of whey is referred to as sweet whey. The second happens in the Cottage cheese manufacture for increased
basic whey type, acid whey, results from processes using firmness) in addition to the acid coagulation.
fermentation or addition of organic or mineral acids to Various technological steps used in the pretreatment of
coagulate the casein as in the manufacture of fresh acid- milk before the main processes (such as various thermal
coagulated cheeses (e.g., Cottage cheese or quark) or most treatments before the casein-clotting operation) may also
industrial acid casein. influence the composition of the whey resulting from such

731
732 Whey Processing | Utilization and Products

Table 1 Typical composition of sweet and acid whey Table 2 Typical composition of major types of dried whey
(g l 1 whey) products (%, w/w)

Component Sweet whey Acid whey Total


Product type protein Lactose Minerals
Total solids 63.0–70.0 63.0–70.0
Lactose 46.0–52.0 44.0–46.0 Regular whey 12.5 73.5 8.5
Protein 6.0–10.0 6.0–8.0 powder
Calcium 0.4–0.6 1.2–1.6 Demineralized 13.7 75.7 3.5
Phosphate 1.0–3.0 2.0–4.5 (70%) whey
Lactate 2.0 6.4 powder
Chloride 1.1 1.1 Demineralized 15.0 83.0 1.0
(90%) whey
Illustrative data compiled from various sources. powder
Ultrafiltration 1.0 90.0 9.0
permeate powder
milk. Typically, the composition of the mineral fraction may Whey protein 35.0 50.0 7.2
be altered slightly and the content of heat-labile whey ‘concentrate’
proteins may be reduced; these changes may result in further (skim milk
replacer)
alterations in the technological properties of such wheys.
Whey protein 65.0–80.0 4.0–21.0 3.0–5.0
New technological alternatives for processing of dairy concentrate
fluids, including membrane processing by ultrafiltration Whey protein isolate 88.0–92.0 <1 2.0–3.0
(UF) of milk in cheese manufacture or fractionation of
Illustrative data compiled from various sources including data from
the various wheys into various whey-based products, pro- manufacturers.
duce a whey-like residue termed UF permeate. The main
difference between UF permeates and the various whey
types is typically the virtual absence of whey proteins from technological properties, depending on various pretreat-
the permeate. Although technically UF permeate does not ment operations used to handle the original milk or the
fit the definition of whey, it is referred to in this article original whey. Some of the typical dried whey products
where appropriate, as its processing and utilization often are listed in Table 2. Partial (70%) or almost complete
present similar challenges and opportunities as for whey. (90%) demineralization of the whey is an important
pretreatment process differentiating many of the whey
powders (see Whey Processing: Demineralization).
Industrial Technologies Used Dried whey powder can be produced also by roller
in the Processing of Whey and UF (drum)-drying; although roller-drying of whey is not
Permeates easy, the process is much cheaper than spray-drying and
the lower quality of the resulting powder may not be
As a general rule, about 9 l of whey is obtained for every detrimental for all applications. Whey, or nowadays
kg of cheese produced; thus, the volume of whey to be more importantly UF permeate from milk or whey, is
processed, originating from just one typical large-scale the principal raw material for the crystallization of
cheesemaking operation, can exceed 1  106 l day 1. lactose (see Lactose and Oligosaccharides:
Most of the technological alternatives used in specialized Lactose: Chemistry, Properties; Lactose: Production,
whey-processing plants are thus large-scale operations, Applications); the residual, partially or more substan-
some with a capacity to handle up to 107 l of whey daily. tially delactosed whey (‘mother liquor’) constitutes yet
The simplest technology for the conversion of whey to another dried whey product differing in composition
industrially valuable products is drying. Typical traditional from the basic dried unmodified whey. Finally, UF
whey-drying operations consist of evaporation in multistage permeates are also being dried with increasing fre-
vacuum evaporators, followed by spray-drying. The equip- quency, giving yet another modified dried whey
ment used does not differ greatly from other such dairy product consisting primarily of lactose and whey miner-
plant installations but the evaporation and drying conditions als but almost devoid of protein. Until recently, spray-
must be adjusted to accommodate the specific properties of drying was the only technique used for the production of
the whey. In particular, the differences between evaporation dried permeates; however, due to the very low market
or spray-drying of skim milk and whey include the need to value of these powders, the spray-drying of permeates is
precrystallize the lactose in whey before the drying step to uneconomical. Several novel technological approaches
minimize the problems of hygroscopicity, as well as careful to drying of milk or whey permeates have been
manipulation of the heat conditions to minimize problems described in the literature and are available to the indus-
related to heat sensitivity of whey proteins. Dried whey try. In principle, these technologies are based on the
powders can differ rather substantially in composition and production of concentrates with very high total solids
Whey Processing | Utilization and Products 733

content (about 76%) combined with additional moisture mind when using dried whey in applications leading to
removal by alternative means, for example, by specially development of supersaturated lactose solutions, for
designed screw conveyors allowing additional evapora- example, the unfrozen fraction of ice cream. The well
tion of water while inducing lactose crystallization. The known – but nowadays rarely seen – sensory defect called
final drying step is accomplished in a fluidized bed, thus sandiness can occur more readily when dried whey is used
avoiding the need for the costly spray-drying. as a portion of nonfat milk solids in ice cream formulation.
The levels and forms of whey proteins are important The various higher value dry whey-based products, listed
factors differentiating the various dried whey protein pro- in Table 2, are increasingly being used as nutritionally
ducts. In contrast to the processes described above, the WPF important components in special dietary products for
can be removed (selectively or totally) from raw whey and infant, geriatric, or sport nutrition. Large amounts of
concentrated by using various membrane processes (see whey are being processed for use in the various infant
Membrane-based fractionation) giving rise to whey protein nutrition products, with demineralized whey, lactose, and
concentrate (WPC), whey protein isolate (WPI), or WPF modified whey protein products being especially important
products (see Milk Protein Products: Whey Protein in this application. Some of the unique technological prop-
Products). Although products with as little as 35% protein erties of whey protein (in particular the acid solubility,
(produced by partial removal of lactose through crystalliza- heat-gelling, and foaming) make the various whey-based
tion or by using simple UF and intended for replacement of high-protein products ideal for use in acid beverages,
skim milk powder in certain applications) are included under foamed dairy desserts, yogurts, and similar dairy and non-
the term WPC, the more valuable WPC products have at dairy products. A heat-denatured whey protein powder
least 65% protein and the production technology does not termed ‘traditional lactalbumin’ has a very high water-
involve any step involving lactose precrystallization. The holding capacity and thus has been shown to be preferable
highest quality WPI and WPF products are manufactured to the undenatured whey protein in applications such as
using various technologies including diafiltration, electro- protein enrichment of pasta dough. In general, it has been
dialysis, ion exchange, nanofiltration, or their combinations. estimated that of about 10 million tonnes of dried whey
For the production of almost all whey protein products, the solids produced annually worldwide, over 50% is being
final step is spray-drying, which should be controlled care- utilized in human foods, with the rest finding less attractive
fully to minimize heat damage of the thermally sensitive valorization in animal feeds.
whey proteins. Both spray-drying and especially evaporation However, liquid unprocessed whey can also be a raw
can cause heat damage resulting in loss of solubility and material for the production of some traditional foods
other functionality defects. Thus, as a preconcentration destined for direct consumer markets. Two such product
step, especially for the production of WPC, WPI, and classes, whey cheeses and whey beverages, represent the
WPF products, reverse osmosis or freeze concentration can traditional uses of whey on a small scale, practiced long
be viewed as viable alternatives to traditional thermal before the industrial approach became feasible, and are
evaporation. The use of reverse osmosis to increase the still important in some parts of the world.
capacity of conventional whey evaporators, or even for pre-
concentrating large amounts of whey before transportation
to a central whey processing facility, is quite common in the Whey Beverages
whey processing industry.
The drinking of whey for therapeutic applications was
already advocated in ancient Greece by Hippocrates.
Utilization of Whey in Industrially This and other similar anecdotal comments concerning
Processed Foods the use of whey as a beverage (including even in the
rhyme Little Miss Muffet. . .) illustrate one of the most
The preceding discussion focused on industrial large-scale obvious, but least industrially advantageous, uses of whey
processes resulting in an array of technological and func- as a drink, logically paralleling the use of milk, buttermilk,
tional whey ingredients or whey protein products, used and other fluid milk products as beverages.
commonly in many processed foods such as spreads, sauces, In general, whey beverages have not been overly suc-
dry soups, beverage and similar mixes, cookies and other cessful with the sophisticated modern consumer, save for a
bakery products, ice cream, and a myriad of other industrial few rather exceptional instances. Minimally processed
food items. Dried whey is a very suitable ‘bulking’ food unflavored or modified raw whey is sold in health-food
ingredient due to its bland taste compatible with many food stores in various countries, especially in Europe, where the
processing applications. The heat sensitivity of whey pro- current organic food movement may result in increased
teins could pose problems in heated liquid food opportunities in this regard. Occasional reports on whey-
applications, but in most solid foods this is of no concern. based beverages marketed for special occasions (e.g., an
Similarly, the limited solubility of lactose must be kept in ‘official Olympic Games drink’ in 1984 in Sarajevo) can
734 Whey Processing | Utilization and Products

be found in the literature. Local markets often feature whey concentrates or as dry powders for home reconstitution
beverages produced by dairy companies looking for new can be occasionally found on regular market shelves,
outlets for their surplus whey; unfortunately, these attempts including even suitably flavored plain dry whey powder.
are typically short-lived, often due to the lack of any serious Various ‘‘miracle protein’’ milk shake formulae based on
product development effort that is necessary preceding the isolated – even hydrolyzed – whey proteins are com-
launch of any such new product. The rather unpleasant monly marketed as products for body builders, active
flavor of raw whey, the origins of which have never been sport enthusiasts, and other health-food applications.
satisfactorily explained, is a major deterrent limiting the
acceptance of these products by contemporary consumers,
especially in view of the fierce competition of other
flavorful and inexpensive thirst-quenching beverages. Whey Cheeses
The only whey beverage with a record of lasting success
is the Swiss product Rivella, which in fact uses highly Whey cheeses have suffered a somewhat similar lack of
modified whey as only a minor ingredient at 33% total international marketing success; however, in a few
volume (the remaining 67% being added water). Other localities, whey cheeses belong among the most tradi-
locally successful products have existed on some European tional and most important foods. Two types of whey
national markets (Austria, Finland, The Netherlands, and cheeses are recognized in the textbooks and by the
Switzerland) for some time; however, several known inter- International Dairy Federation. The main differences
national marketing attempts with these products have failed. between the production technologies for these two pro-
The most typical approach to whey beverage development ducts are illustrated schematically in Figure 1. The
is combination with fruit juices, especially citrus fruits, more widespread of the two is the Italian-type whey
which are most compatible with the flavor characteristic of cheese Ricotta; similar products are also popular in
whey. This is especially true in the case of acid whey, which Portugal, Turkey, and other localities. These products
is more suitable for this application due to its high content of are essentially a heat–acid-coagulated whey protein
lactic acid and calcium in comparison to sweet whey. Some paste, sometimes referred to as ‘whey quark’
of the fruit flavors used in commercial whey beverages (see Cheese: Acid- and Acid/Heat Coagulated
include mango, passion fruit, grapefruit, lemon, orange, Cheese). The processing technology is quite simple,
pear, or their combinations. Other approaches documented consisting of heating whey (often mixed with up to
in the literature include the production of yogurt drinks 25% added skim milk) to at least 90  C for a few
containing a substantial whey component, or fermentation minutes, resulting in heat-induced coagulation of whey
of liquid WPCs from sweet whey to produce a ‘thin sour proteins (and any caseins if present due to the added
milk’, as the stability of the whey protein at low pH does not milk); the coagulum is then separated by suitable
lead to the clotted appearance of traditional sour milk mechanical means. The traditional batch manufacturing
products. Milk or whey UF permeates are suitable for the procedures are cumbersome and involve much
production of isotonic sport drinks or beverages destined for
replenishment of the mineral balance after vigorous physical
activity; several such products have been described in
commercial product surveys.
Attempts to produce whey beverages with high whey
protein content have been recorded in the literature and
are often seen in various food trade exhibitions, with little
or no indication of commercial viability. It may be that
with the presently increasing reputation of whey proteins
as nutraceutically important food components, the devel-
opment of such products will be intensified. The heat
sensitivity of whey proteins is one important problem
encountered in manufacturing these products with a
demand for extended shelf life. However, since the resis-
tance of whey proteins to heat-induced coagulation in the
absence of casein increases dramatically at pH below 3.9,
it may be possible to formulate high whey protein drinks,
even for ultra-high temperature (UHT) processing.
Alternatively, various nonthermal processes now being
studied actively for various food processing uses can be
considered. Whey beverages offered as liquid Figure 1 Alternative processes for manufacturing whey cheeses.
Whey Processing | Utilization and Products 735

hand labor, in particular in the separation of the heat- taste, intensity of the brown color development, and
coagulated, fragile whey protein curd. Mechanized and sometimes inclusion of sweetening agents or other
automated continuous systems are now in existence and nondairy ingredients such as hazelnuts or chocolate.
these result in increased economy, improved shelf life, Because the relatively simple Mysost process technol-
and better sensory qualities of the final product. ogy leaves no residue other than the evaporated water, its
In contrast to the Ricotta-type whey cheese involving principle is much more useful than that of the Ricotta
primarily the protein fraction of whey (and thus not offer- process as a means of whey disposal, particularly for small
ing a solution to the whey disposal problem, which is cheese manufacturers. However, significant product
especially pressing for the small-scale cheese manufac- development efforts would be necessary to modify the
turers), the Norwegian-style whey cheese Mysost utilizes sensory profiles of the basic Norwegian products to suit
all the whey components and leaves no residue other than the tastes of markets outside Norway, as demonstrated in
water vapor. The principle of the Mysost cheese process is several consumer studies conducted in Canada and
even simpler than that for the manufacture of Ricotta, as elsewhere.
Mysost is essentially highly concentrated whey to which
some other components (such as milk fat, cream, or goats’
milk in the most traditional version) have been added
(Table 3). The main technological problem encountered Utilization of Whey as a Fermentation
in the manufacture of Mysost cheese is the crystallization Substrate for Food or Nonfood
of lactose in the highly concentrated whey-based paste Applications
related to the low solubility of lactose. Thus, a ‘controlled
crystallization’ step, consisting of rapid cooling of the hot Although whey and whey-like UF permeates originate
paste with intensive stirring, from about 95  C (the tem- from milk and contain about 50% of valuable milk
perature of the last phase of the evaporation process) to nutrients, the profitable utilization of these materials
for human nutrition continues to be a problem because
below 65  C, is the most essential aspect of the otherwise
of the sheer volume of these by-products of conven-
routine evaporation process carried out in two steps. After
tional or modern cheesemaking. In numerous research
preconcentration in traditional dairy evaporators, the final
reports and several major industrial projects, whey – due
operation involves a special kettle, ‘gryta’, in which batch
to its suitable content of a fermentable carbohydrate,
evaporation is carried out at an elevated temperature
lactose – was used as a medium for the production of
needed to reduce the viscosity of the thick paste. This
various food-grade or nonfood products using microbial
high temperature tends to minimize the uncontrolled
fermentations. Among the most successful current uses
crystallization of lactose and is the main reason for the
of whey in this regard are the plants producing food-
typical brown color of the product due to the pronounced grade or industrial alcohol in New Zealand, the United
Maillard reaction. The subsequent rapid cooling step in States, Ireland, and possibly other countries; one of the
scraped-surface agitators/coolers promotes formation of best examples of such use is the Original Bailey’s Cream
very small lactose crystals, thus minimizing the develop- Liqueur from Ireland or most of the liqueurs produced
ment of pronounced grittiness/sandiness of the final in New Zealand. In the past, whey was an important
product. There are two basic types of Mysost – sliceable substrate for conversion into Torula spp. yeast biomass
or spreadable – available in Norway, its country of origin used in animal feeds, or for other fermentation-based,
and still the only significant market for these products; the lactose-derived products such as antibiotics, lactic acid,
two main types are differentiated principally by the moist- or other microbial metabolites. The current preoccupa-
ure content and encompass further variations differing in tion with organically produced foods and natural food
the fat content, inclusion of some goats’ milk for a stronger ingredients may signal a possible opportunity for revi-
talization of some of these processes, abandoned in the
past due to unfavorable economic feasibility in compar-
Table 3 Compositional characteristics of whey cheeses
(%, w/w) ison to direct chemical synthesis or using other
fermentable substrates. Use of whey for the propagation
Cheese Moisture Fat Protein Lactose of lactic cultures for cheese manufacturing is well estab-
Ricotta lished in many countries. Whey can be used also for the
Whole milk 72.2 12.7 11.2 3.0 generation of biogas; several such installations where
Whey 82.5 0.5 11.3 1.5 biogas is used as an energy source in the same cheese
Mysost plant at which whey is produced are in operation in
Sliceable 17.4 28.3 11.5 36.2
Switzerland and possibly other countries. More recently,
Spread 26.6 3.6 7.7 46.2
whey or whey permeates have been shown to be suitable
Illustrative data compiled from various sources. fermentation substrates for the production of
736 Whey Processing | Utilization and Products

bacteriocins such as nisin, or for other valuable food glucose and galactose. Most of the industrial lactose
ingredients using specific strains of various lactic acid hydrolysis processes developed for the production of
bacteria. sweetening syrups for food uses (such as in ice cream)
Lactose is not fermented by many microorganisms, using immobilized -galactosidase enzyme reactors
necessitating careful selection of the cultures for many have failed mainly because of the high cost of such
of the possible fermentation applications. Uses of whey as technology. Alternative low-cost processes, based, for
a raw material for the production of alcoholic beverages, example, on the use of mechanically disrupted
such as whey wine or whey beer, have been described in common dairy bacteria producing high amounts of
the literature, but because lactose is not fermented by -galactosidase, or on other principles, continue to be
common yeasts without special pretreatments, as well as investigated. In general, the lactose hydrolysis process,
other reasons, these attempts at profitable whey utiliza- which seems to be grossly underutilized by the indus-
tion have not resulted in lasting success. However, since try today, could have a major impact on expanding the
the key to new approaches to utilization of whey is to find dairy markets for lactose- intolerant consumers not
new avenues for conversion or other utilization applica- only for the processing and utilization of whey, but
tions of lactose, the fermentation route will undoubtedly also for uses in other dairy and especially nondairy
continue to be explored. One of the currently active foods (see Enzymes Exogenous to Milk in Dairy
industrial research topics is the conversion of lactose to Technology: -D-Galactosidase, Lactose and
oligosaccharides, either by fermentation with whole Oligosaccharides: Lactose: Chemistry, Properties;
microorganisms or by applications of crude enzymatic Lactose: Production, Applications).
extracts of suitable bacterial cultures. Similarly, produc- The main focus of interest regarding the nutritional/
tion of exopolysaccharides from lactose is increasingly nutraceutical properties of whey is currently centered
being used in the production of yogurt; it is likely that on the WPF. Today, whey protein products are the main
this route could be explored in large-scale whey fermen- ingredients in most protein formula supplements used
tations as well.
by bodybuilders and active athletes worldwide. Whey
protein products have been one of the principal ingre-
dients in infant feeding formulae for some time, often
Nutritional and Nutraceutical Aspects
requiring additional treatments such as enzymatic
of Whey Utilization
hydrolysis to minimize the allergenicity potential of
some of the components such as -lactoglobulin. Most
The new approach to developing technically and eco-
recently, whey protein and/or its main fractions have
nomically feasible uses of whey and whey-like products
been investigated for many unique health-enhancing or
lies in finding some unique properties of at least some
disease-combating properties, including immunopoten-
of the main whey components. The current trend in the
food field, focusing on the health-promoting aspects of tiation through increased intracellular glutathione
traditional or novel foods, has opened up new possibi- synthesis; role in reducing cancer cell proliferation;
lities for whey-based products. Whey contains many counteracting the wasting syndrome in HIV-positive
minor milk components that are known (or thought) individuals; antimicrobial properties of some of the
to have physiologically important functions. Some of minor whey proteins; and other effects. WPIs advertised
these compounds are found especially among whey to supply some of these benefits are now being marketed
proteins; these include minor whey protein components even as dietary supplements at highly inflated prices.
such as lactoferrin or lactoperoxidase, the immunoglo- Hydrolysates of whey proteins have been shown to con-
bulins, and even major constituents of the WPF such as tain bioactive peptides effective in blood pressure
the GMP or bovine serum albumin. Some of the reduction, and several industrial products are now on
mineral compounds, especially the calcium phosphate the market.
complex, are now also being marketed as ‘natural’ milk- Extraction of some functional components from whey
based food ingredients. Even lactose is being has become a major industrial activity for specialized
reexamined for its unique nutritional properties, whey processors, although in most cases, the physiologi-
including the purported enhancement of calcium cal functionality in human subjects still awaits
absorption or its unique disaccharide composition, confirmation from well-controlled clinical trials. Of
thus serving as a raw material for the production of course, manufacture of any such isolated functional pro-
prebiotic compounds such as galacto-oligosaccharides duct from the liquid whey will not alleviate requirements
(see Lactose and Oligosaccharides: Lactose: Galacto- for further processing of the residual by-product. As many
Oligosaccharides) or heterooligosaccharides. The lim- of the components with documented or proposed unique
ited sweetness of lactose can be enhanced by functionalities are present in the whey in very minute
hydrolysis into the two lactose monosaccharides, amounts, the need to process the bulk of the residual
Whey Processing | Utilization and Products 737

whey stream will remain a major challenge accompanying See also: Cheese: Acid- and Acid/Heat Coagulated
any such novel development. Cheese. Enzymes Exogenous to Milk in Dairy
Surprisingly, by far the most abundant whey compo- Technology: -D-Galactosidase. Lactose and
nent, water, seems to have been neglected in the Oligosaccharides: Lactose: Chemistry, Properties;
innumerable research attempts to find new profitable Lactose: Galacto-Oligosaccharides; Lactose:
uses of whey and/or its components. The only well- Production, Applications. Milk Protein Products:
known use of whey water is as a condensate from the Membrane-Based Fractionation; Whey Protein Products.
evaporators, mainly for rinsing and equipment washing Whey Processing: Demineralization.
purposes, but not for productive economically advanta-
geous applications. As one potential such use suggested in
the past, the adjustment of the mineral content of the Further Reading
whey water by suitable membrane processing could lead
to various pharmaceutical uses for cleansing solutions Anonymous (2001) The importance of whey and whey components in
food and nutrition. Proceedings of the International Whey Conference,
mimicking the human body fluids. 411pp. Munich, Germany. Hamburg, Germany: B. Behr’s Verlag.
The approach to whey processing and utilization IDF (1998) Whey: Proceedings of the 2nd International Whey
has changed in the recent past, from considering whey Conference, 367pp. Chicago, IL, USA, 27–29 October 1997.
Brussels, Belgium: International Dairy Federation.
a bothersome waste to capitalizing on the opportu- Jelen P (1999) Whey: Composition, properties, processing, and uses.
nities that the whey and whey-like products present In: Francis FJ (ed.) Encyclopedia of Food Science and Technology,
for the innovative technically advanced processor. 2nd edn., Vol. 4, pp. 2652–2661. New York: John Wiley.
Jelen P (2009) Whey-based functional beverages. In: Paquin P (ed.)
However, for traditional cheese manufacturers, the Functional and Speciality Beverage Technology, pp. 259–280.
problems of dealing with large volumes of whey or Cambridge, UK: Woodhead Publishing Ltd.
whey permeate continue to pose difficulties, determin- Modler HW (2000) Milk processing. In: Nakai S and Modler HW (eds.) Food
Proteins: Processing Applications, pp. 1–88. New York: Wiley-VCH.
ing the overall success of their main industrial activity. Onwulata C and Huth P (2008) Whey Processing, Functionality and
Nowadays, industrial processing of whey is a highly Health Benefits, 416pp. Hoboken, NJ: Wiley-Blackwell.
specialized, technologically advanced segment of the Sienkiewicz T and Riedel C-L (1990) Whey and Whey Utilization, 379pp.
Gelsenkirchen, Germany: Verlag Th. Mann.
dairy industry requiring up-to-date knowledge and Zadow JG (ed.) (1992) Whey and Lactose Processing, 489pp. London:
focused attention. Elsevier Applied Science.

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