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Zootaxa 1579: 41–53 (2007) ISSN 1175-5326 (print edition)

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ZOOTAXA
ISSN 1175-5334 (online edition)

Notes to the current checklist of Tardigrada

PETER DEGMA1 & ROBERTO GUIDETTI2

1
Department of Zoology, Comenius University in Bratislava, Mlynská dolina B-1, SK-84215Bratislava, Slovakia.
E-mail: degma@fns.uniba.sk
2
Department of Animal Biology, University of Modena and Reggio Emilia, Modena, via Campi 213/d, 41100, Modena, Italy.
E-mail: guidetti.roberto@unimore.it

Abstract

Corrections and additions to the recently published checklist of Tardigrada taxa (Guidetti & Bertolani 2005) are defined
and described until the acceptance of this article. Two new synonyms are stated here: Hypsibius iharosi Bartoš, 1941 syn.
nov. and Isohypsibius gibbus (Marcus, 1928) syn. nov.

Key words: Tardigrada, checklist, Hypsibius iharosi syn. nov., Isohypsibius gibbus syn. nov.

Introduction

Until recently, only two papers dealing with worldwide tardigrade taxa (i.e. the monograph of Ramazzotti &
Maucci 1983, and the checklist of limno-terrestrial species of McInnes 1994) were available for specialists of
this group and for all researchers interested in biodiversity. In the period after these papers were published the
number of described taxa had dramatically increased, along with several changes to the taxonomic position of
other species, which called for the recent publication by Guidetti & Bertolani (2005) to draw this information
together. After this publication we noted that a revision of the checklist was needed due mainly to typing
errors, the lack of inclusion of all species, incorrectness in name spellings or years of publication and an
incomplete reference list. During the review process, we thoroughly applied rules and recommendations of
International Code of Zoological Nomenclature — Fourth Edition (1999) (especially Article 22:
Recommendations 22A.2.2., 22A.3; Article 51: Recommendations 51A and 51B). We also used parentheses
for author and year in cases when a subgenus has been elevated to genus level (Article 51.3; Alessandro
Minelli pers. com.). Some of the taxa were supplemented by new species and by bibliographic references of
inherent synonymies and/or the amended diagnoses not described prior to or not mentioned in Guidetti &
Bertolani (2005).
We report all differences between the published checklist (Guidetti & Bertolani 2005) and the revised
checklist in Table 1. In addition, some new synonyms are reported.

Acknowledgements

We would like to thank Clark W. Beasley (U.S.A.), Roberto Bertolani (Italy), Hieronymus Dastych (Ger-
many), Paolo Fontoura (Portugal), Joel Hallan (U.S.A.), Łukasz Kaczmarek (Poland), DeeDee Kathman
(U.S.A.), Sandra J. McInnes (U.K.), Łukasz Michalczyk (U.K.), Giovanni Pilato (Italy) and Alessandro

Accepted by C. Beasley: 28 Jul. 2007; published: 10 Sept. 2007 41

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Minelli former president of International Commission on Zoological Nomenclature (Italy) for consultations
and suggestions. We thank also to Clark W. Beasley for the English revision of the paper and to two anony-
mous reviewers for their valuable comments.
This study is a component of the implementation of a Project No. 1/3265/06 supported by the Slovak Sci-
entific Grant Agency (VEGA). Financial support for the research to R. Guidetti came from FAR ex 60%
MURST grant.

TABLE 1. Corrections and additions to the checklist (and related references) of Tardigrada (Guidetti & Bertolani 2005);
page numbers in the table refer to that paper. The assignment of synonyms and species inquirenda are reported within
square brackets‘ [ ] ’ (if the reference is not specified, then the assignments were already reported by Ramazzotti &
Maucci 1983). Amendments are reported within angle brackets ‘ < > ’. Comments are reported within brace parentheses
‘ { } ’. The species written in bold characters are the genus type species, those preceded by asterisk ‘ * ’ were described
or amended after or were not included in Guidetti & Bertolani (2005). The references preceded by asterisk ‘ * ’ were not
included in the paper of Guidetti & Bertolani (2005), those not reported in this table are present in that paper.

p. 5
*Bartoš, E. (1941) Studien über die Tardigraden des Karpathengebietes. Zoologischer Jahrbücher, Abteilung für System-
atik, Ökologie und Geographie der Tiere, 74, 435–472.
*Beasley, C.W., Kaczmarek, Ł. & Michalczyk, Ł. (2006) Redescription of Doryphoribius vietnamensis (Iharos, 1969)
(Tardigrada) comb. nov. on the basis of the holotype and additional material from China. Acta Zoologica Academiae
Scientiarum Hungaricae, 52, 367–372.

p. 6
*Binda, M.G., Pilato, G., Moncada, E. & Napolitano, A. (2001) Some tardigrades from Central Africa with the descrip-
tion of two new species: Macrobiotus ragonesei and M. priviterae (Eutardigrada Macrobiotidae). Tropical Zoology,
14, 233–242.
*Biserov, V.I. (1986) Terrestrial water bears from the north Caucasus. Zoologicheskii Zhurnal, 65, 741–756. (in Russian)
*Cuénot, L. (1932) Tardigrades. Faune de France, 24, 1–96. Ed. Paul Lechevalier, Paris.
Dastych, H. (1973a) Systematic studies on Tardigrada, II. Echiniscus clavisetosus Mihelčič, 1958, a synonym of Echinis-
cus spitzbergensis Scourfield 1897. Bulletin de la Société des Amis des Sciences et des Lettres de Poznań, 14, 89–99.
*Dastych, H. (1973b) Redescription of Macrobiotus spectabilis Thulin, 1928 (Tardigrada). Bulletin de l’Academie
Polonaise des Sciences, Série des sciences biologiques CI, II, XXI (12), 823–825.
*Dastych, H. (1983) Two new Eutardigrada species from west Spitsbergen and the Tatra Mts. Bulletin de la Société des
Amis des Sciences et des Lettres de Poznań, Serie D, sciences biologiques, 23, 195–200.

p. 7
*Dastych, H. (1992) Paradiphascon manningi gen. n. sp. n., a new water-bear from South Africa, with the erecting of a
new subfamily Diphasconinae (Tardigrada). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Insti-
tut, 89, 125–139.
Dastych, H. (2001) Notes on the revision of the genus Mopsechiniscus (Tardigrada). Zoologischer Anzeiger, 240, 299–
308.
Dastych, H., Kraus, H. & Thaler, K. (2003) Redescription and notes on the biology of the glacier tardigrade Hypsibius
klebelsbergi Mihelčič, 1959 (Tardigrada), based on material from the Ötztal Alps, Austria. Mitteilungen aus dem
Hamburgischen Zoologischen Museum und Institut, 100, 73–100.
Dastych, H., Ryan, P.G. & Watkins, B.P. (1990) Notes on Tardigrada from western Dronning Maud Land (Antarctica)
with a description of two new species. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 10,
57–66.
Dastych, H. & Thaler, K. (2002) The tardigrade Hebesuncus conjungens (Thulin, 1911) in the Alps, with notes on mor-
phology and distribution (Tardigrada). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14,
83–94.
de Zio Grimaldi, S., D’Addabbo Gallo, M., Morone De Lucia, R.M. & Troccoli, A. (1995/96) Hemitanarctus chimaera
n.g., n.sp., new Halechiniscidae from the Ionian Sea (Tardigrada, Heterotardigrada). Zoologischer Anzeiger, 234,
167–174.

42 · Zootaxa 1579 © 2007 Magnolia Press DEGMA & GUIDETTI

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p. 8
Gallo D’Addabbo, M., de Zio Grimaldi, S. & Sandulli, R. (2001) Heterotardigrada of two submarine caves in S. Domino
Island (Tremiti Island) in the Mediterranean Sea, with the description of two new species of Stygarctidae. Zoologis-
cher Anzeiger, 240, 361–369.

p. 9
*Guidetti, R., Bertolani, R. & Degma, P. (2007) New taxonomic position of several Macrobiotus species (Eutardigrada:
Macrobiotidae). Zootaxa, 1471, 61–68.
*Guidetti, R., Gandolfi, A., Rossi, V. & Bertolani, R. (2005) Phylogenetic analysis of Macrobiotidae (Eutardigrada,
Parachela): a combined morphological and molecular approach. Zoologica Scripta, 34, 235–244.
Guidetti, R. & Pilato, G. (2003) Revision of the genus Pseudodiphascon (Tardigrada, Macrobiotidae), with the erection
of three new genera. Journal of Natural History, 37, 1679–1690.
Hansen, J.G., Gallo D’Addabbo, M. & de Zio Grimaldi, S. (2003) A comparison of morphological characters within the
genus Rhomboarctus (Tardigrada: Heterotardigrada) with the description of two new species. Zoologischer
Anzeiger, 242, 83–96.
*Kaczmarek, Ł., Beasley, C.W. & Michalczyk, Ł. (2006) The first record of the genus Haplohexapodibius Pilato & Bea-
sley, 1987 in Africa, with notes on synonymy of Hexapodibius beasleyi Maucci, 1988, with Haplohexapodibius
seductor Pilato & Beasley, 1987. African Zoology, 41, 290–293.
*Kaczmarek, Ł. & Michalczyk, Ł. (2006a) Thulinius itoi comb. nov. a new systematic position for Isohypsibius itoi (Tsu-
rusaki, 1980) (Tardigrada: Eutardigrada: Hypsibiidae). Zootaxa, 1373, 65–68.
*Kaczmarek, Ł. & Michalczyk, Ł. (2006b) Redescription of Macrobiotus tridigitus Schuster, 1983 and the erection of a
new genus of Tardigrada (Eutardigrada, Macrobiotidae). Journal of Natural History, 40, 1223–1229.
Kristensen, R.M. (1987) Generic revision of the Echiniscidae (Heterotardigrada), with a discussion of the origin of the
family. In: R. Bertolani (ed.), Biology of Tardigrades, Sel. Symp. Monogr. U.Z.I., 1, Mucchi, Modena, 261–335.
*Iharos, A. (1940) Additions to the Hungarian fauna of tardigrades. A Keszthelyi Premontrei Gimnázium Évkönyve, 12,
1–20. (in Hungarian)
*Marcus, E. (1928) Bärtierchen (Tardigrada). Die Tierwelt Deutschlands, Verlag von Gustav Fischer, Jena, 12 (IV), I–
VII + 1–230.
*Maucci, W. (1981) Hexapodibius castrii, nuova posizione sistematica per Hypsibius (Calohypsibius) castrii Ramaz-
zotti, e considerazioni sul genere Hexapodibius Pilato (Eutardigrada, Hypsibiidae). Atti Società italiana di Scienze
naturali Museo civico di Storia naturale Milano, 122 (1-2), 32–36.
*Maucci, W. (1985) Materiali per una revisione del genere Echiniscus Schultze, 1840. I. Il complesso blumi (Heterotar-
digrada, Echiniscidae). Bollettino del Museo civico di Storia Naturale, Verona, 12, 109–139.
*Maucci, W. (1986) Tardigrada. In Fauna d’Italia. Vol 24. Edizioni Calderini, Bologna, pp. 388.
*Maucci, W. & Pilato, G. (1974) Macrobiotus spectabilis Thulin, 1928 e Macrobiotus grandis Richters, 1911: due buone
specie di eutardigradi. Animalia, 1(1/3), 245–255.

p. 10
*Michalczyk, Ł. & Kaczmarek, Ł. (2006) Revision of the Echiniscus bigranulatus group with a description of a new spe-
cies Echiniscus madonnae (Tardigrada: Heterotardigrada: Echiniscidae) from South America. Zootaxa, 1154, 1–26.
*Mihelčič, F. (1938a) Beiträge zur Kenntnis der Tardigrada Jugoslawiens. Zoologischer Anzeiger, 122, 318–321.
*Mihelčič, F. (1938b) Beiträge zur Kenntnis der Tardigrada Jugoslawiens. II. Zoologischer Anzeiger, 121, 349–350.
*Mihelčič, F. (1951) Beitrag zur Systematik der Tardigraden. Archivio Zoologico Italiano, 36, 57–103.
*Mihelčič, F. (1953) Contribucion al conocimiento de los tardigrados con especial consideracion de los tardigrados de
Osttirol. Anales de edafologia y fisiologia vegetal, 12, 431–479.
*Mihelčič, F. (1964) Eine neue Echiniscus-Art (Tardigrada) aus dem Karst (Echiniscus (E.) inocellatus n.sp.). Zoologis-
cher Anzeiger, 172, 239–242.
*Murray, J. (1908) Encystment of Tardigrada. Transactions of the Royal Society of Edinburgh, 45(4) [issued also sepa-
rately in 1907], 837–854.
*Petersen, B. (1951) The tardigrade fauna of Greenland, a faunistic study with some few ecological remarks. Med-
delelser om Grønland, 150 (5), 1–94.
*Pilato, G. (1969c). Su alcuni tardigradi muscicoli delle Alpi Apuane. Bollettino della Accademia Gioenia di Scienze
Naturali, Catania, Serie IV, 10, 194–204.
*Pilato, G. (1973c) Redescription of Haplomacrobiotus hermosillensis May, 1948, and consideration on the genus Hap-
lomacrobiotus (Eutardigrada). Zeitschrift für Zoologische Systematik und Evolutionsforschung, Hamburg, 11 (4),
283–286.
*Pilato, G. (1984) Sul valore di buona specie di Macrobiotus mandalaae Pilato, 1974 (Eutardigrada). Animalia, 11, 17–
21.
*Pilato, G. (2006) Remarks on the Macrobiotus polyopus group, with the description of two new species (Eutardigrada,

NOTES TO THE CURRENT CHECKLIST OF TARDIGRADA Zootaxa 1579 © 2007 Magnolia Press · 43
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Macrobiotidae). Zootaxa, 1298, 37–47.

*Pilato, G., Guidetti, R., Rebecchi, L., Lisi, O., Hansen, J.G. & Bertolani, R. (2006) Geonemy, ecology, reproductive
biology and morphology of the tardigrade Hypsibius zetlandicus (Eutardigrada: Hypsibiidae) with erection of Bore-
alibius gen. n. Polar Biology, 29, 595–603.

p. 12
*Rho, H.S., Moon, S.Y., Chang, C.Y. & Kim, W. (1997) Four freshwater eutardigrades from Korea with description of
Isohypsibius brevitubulatus n. sp. Korean Journal of Biological Science, 1, 271–279.
*Thulin, G. (1911) Beiträge zur Kenntnis der Tardigradenfauna Schwedens. Arkiv för Zoologi, 7 (16), 1–60.
*Van Rompu, E.A., De Smet, W.H. & Coomans, A. (1995) Some terrestrial tardigrades from Zimbabwe. Biologisch-
Jaarboek, 62, 48–55.
Villora-Moreno, S. & de Zio Grimaldi, S. (1993) Redescription and ecology of Batillipes phreaticus Renaud-Debyser,
1959 (Arthrotardigrada, Batillipedidae) in the gulf of Valencia (Western Mediterranean). Cahiers de Biologie
Marine, 34, 387–399.

p. 13
*Batillipes spinicauda Gallo D’Addabbo, Sandulli & de Zio Grimaldi, 2005

p. 14
*Coronarctus verrucatus Hansen, 2007

Dipodarctus borrori Pollock, 1995 [Hemitanarctus chimaera de Zio Grimaldi, D'Addabbo Gallo, Morone De Lucia &
Troccoli, 1995/96 in de Zio Grimaldi et al. 1995/96]

p. 15
*Parmursa torquata Hansen, 2007

p. 16
*Halechiniscus jejuensis Chang & Rho, 2002
Halechiniscus remanei remanei Schulz, 1955
*Halechiniscus remanei antillensis Renaud-Mornant, 1985

Orzeliscus du Bois-Reymond Marcus, 1952

Orzeliscus belopus du Bois-Reymond Marcus, 1952

p. 17
Rhomboarctus aslaki Hansen, Gallo D’Addabbo & de Zio Grimaldi, 2003
Rhomboarctus duplicicaudatus Hansen, Gallo D’Addabbo & de Zio Grimaldi, 2003

Styraconyx hallasi Kristensen, 1977 <Amended by Kristensen & Higgins 1984>

Styraconyx haploceros Thulin, 1942 <Amended by Kristensen & Higgins 1984>

Tetrakentron Cuénot, 1892 {corrected according to Cuénot 1932}

Tetrakentron synaptae Cuénot, 1892 {corrected according to Cuénot 1932}

p. 18
Neoarctidae (de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992)
<Amended by Bello & de Zio Grimaldi 1998>

Neoarctus primigenius de Zio Grimaldi, D'Addabbo Gallo & Morone De Lucia, 1992

Neostygarctidae de Zio Grimaldi, D’Addabbo Gallo & De Lucia Morone, 1987

<Amended by Bello & de Zio Grimaldi 1998; de Zio Grimaldi et al. 1998>

p. 19
*Megastygarctides christinae Hansen & Kristensen, 2006
*Megastygarctides gerdae Hansen & Kristensen, 2006

Pseudostygarctus intermedius (Renaud-Mornant, 1979) {formerly Mesostygarctus intermedius} <Amended by Bello &
de Zio Grimaldi 1998>

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Stygarctus gourbaultae Renaud-Mornant, 1981

p. 20
Echiniscoides Plate, 1888 {corrected according to Marcus 1936}
<Amended by Kristensen & Hallas 1980>

Oreella mollis Murray, 1910 {egg described in Bertolani et al. 1996} [Oreella minor Ramazzotti, 1964 in Dastych et al.
1998] <Amended by Dastych et al. 1998>

p. 21
Bryochoerus intermedius hawaiicus (Thulin, 1928)
Bryochoerus intermedius laevis (Marcus, 1936)

Bryodelphax alzirae (du Bois-Reymond Marcus, 1944)

Bryodelphax amphoterus (Durante Pasa & Maucci, 1975)
*Bryodelphax asiaticus Kaczmarek & Michalczyk, 2004
*Bryodelphax brevidentatus Kaczmarek, Michalczyk & Degma, 2005
Bryodelphax dominicanus (Schuster & Toftner, 1982)
Bryodelphax iohannis Bertolani, Guidi & Rebecchi, 1996
*Bryodelphax lijiangensis Yang, 2002
Bryodelphax mateusi (Fontoura, 1982)
Bryodelphax ortholineatus (Bartoš, 1963)
Bryodelphax sinensis (Pilato, 1974)
Bryodelphax tatrensis (Węglarska, 1959)
Bryodelphax weglarskae (Pilato, 1972)

Cornechiniscus ceratophorus (Maucci, 1973)

Cornechiniscus cornutus (Richters, 1906) [Pseudechiniscus intermedius Mihelčič, 1970]
Cornechiniscus madagascariensis Maucci, 1993

p. 22
*Echiniscus arthuri Pilato, Binda & Lisi, 2005
Echiniscus batramiae Iharos, 1936
*Echiniscus bigranulatus Richters, 1907 <Amended by Michalczyk & Kaczmarek 2006>
Echiniscus bisetosus Heinis, 1908 {not valid according to Maucci 1985}
Echiniscus blumi blumi Richters, 1903 [Echiniscus ramazzottii Binda & Pilato, 1969 in Maucci 1985]
Echiniscus blumi schizofilus Bartoš, 1941
Echiniscus dearmatus Bartoš, 1935 [Echiniscus szaboi Iharos, 1973]
Echiniscus dikenli Maucci, 1973
Echiniscus dominicanus Schuster & Toftner, 1982 see Bryodelphax dominicanus
Echiniscus dreyfusi de Barros, 1942
*Echiniscus elaeinae Pilato, Binda & Lisi, 2005
Echiniscus evelinae de Barros, 1942
Echiniscus filamentosus Plate, 1888
*Echiniscus ganczareki Michalczyk & Kaczmarek, 2007

p. 23
Echiniscus granulatus (Doyère, 1840) [Echiniscus crassus Richters, 1904; Echiniscus fortis Bartoš, 1935; Echiniscus
abanti Maucci, 1973]
*Echiniscus hexacanthus Maucci, 1973
Echiniscus inocelatus Mihelčič, 1938 [Echiniscus inocellatus Mihelčič, 1964] {this species was first described as Echi-
niscus granulatus inocelatus in Mihelčič (1938a), then in 1964, Mihelčič described this species based on different
type material (due to the loss of the original material) but stressing the same characters, elevating its taxonomical
rank and changing its name in Echiniscus inocellatus, therefore we consider E. inocellatus as synonymous}
Echiniscus inocellatus Mihelčič, 1964 see Echiniscus inocelatus
*Echiniscus madonnae Michalczyk & Kaczmarek, 2006
Echiniscus marinellae Bartoš, 1935 see Testechiniscus spitsbergensis
Echiniscus melanophthalmus Bartoš, 1936 see Testechiniscus spitsbergensis
Echiniscus menzeli Heinis, 1917 see Testechiniscus spitsbergensis

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Echiniscus merokensis merokensis Richters, 1904 [Echiniscus iharosi Rudescu, 1964]

Echiniscus militaris Murray, 1911 {Echiniscus militaris hystrix Maucci, 1973-74 included in Ramazzotti & Maucci 1983
is here omitted as not valid according to Maucci 1985}
*Echiniscus nelsonae Li, Wang & Yu, 2007

p. 24
*Echiniscus ollantaytamboensis Nickel, Miller & Marley, 2001 <Amended by Michalczyk & Kaczmarek 2006>
Echiniscus phocae du Bois-Reymond Marcus, 1944
Echiniscus quadrispinosus quadrispinosus Richters, 1902 [Echiniscus scrofa Richters, 1902]
*Echiniscus quitensis Pilato, 2007
Echiniscus ramazzottii Binda & Pilato, 1969 see Echiniscus blumi
*Echiniscus ranzii Ramazzotti, 1964 <Amended by Michalczyk & Kaczmarek 2006>
Echiniscus rosaliae Mihelčič, 1951 see Testechiniscus spitsbergensis
Echiniscus rufoviridis du Bois-Reymond Marcus, 1944
*Echiniscus shaanxiensis Li, Wang & Yu, 2007
*Echiniscus taibaiensis Wang & Li, 2005
Echiniscus tamus Mehlen, 1969
Echiniscus testudo (Doyère, 1840) [Echiniscus bellermanni C.A.S. Schultze, 1840; Echiniscus inermis Richters, 1902;
Echiniscus trifilis Rahm, 1921; Echiniscus filamentosus mongoliensis Iharos, 1973]

p. 25
Echiniscus trojanus Maucci, 1973
*Echiniscus viridianus Pilato, Fontoura & Lisi, 2007
Echiniscus viridissimus Péterfi, 1956

Hypechiniscus gladiator gladiator (Murray, 1905) [Parechiniscus unispinosus da Cunha, 1947]

Hypechiniscus gladiator bigladii (Iharos, 1973)
Hypechiniscus gladiator exarmatus (Murray, 1907) see Hypechiniscus exarmatus
Hypechiniscus gladiator fissigladii (Iharos, 1973)
Hypechiniscus gladiator spinulosa (Iharos, 1973)

Mopsechiniscus du Bois-Reymond Marcus, 1944

<Amended by Kristensen 1987; Dastych 2001>

Novechiniscus armadilloides (R.O. Schuster, 1975)

p. 26
Pseudechiniscus bartkei Węglarska, 1962
*Pseudechiniscus beasleyi Li, Wang & Yu, 2007
Pseudechiniscus facettalis Petersen, 1951 [Pseudechiniscus pseudoconifer facettalis Maucci, 1954]
*Pseudechiniscus gullii Pilato & Lisi, 2006
Pseudechiniscus insolitus Maucci, 1991
Pseudechiniscus juanitae de Barros, 1939 [Pseudechiniscus suillus franciscae de Barros, 1942]
Pseudechiniscus novaezeelandiae novaezeelandiae (Richters, 1908)
Pseudechiniscus novaezeelandiae aspinosa Iharos, 1963
Pseudechiniscus novaezeelandiae laterospinosa Iharos, 1963
Pseudechiniscus novaezeelandiae marinae Bartoš, 1934
*Pseudechiniscus papillosus Li, Wang, Liu & Su, 2005
Pseudechiniscus ramazzottii facettalis Iharos, 1964 [Pseudechiniscus ramazzottii lineatus Maucci, 1973-74]
Pseudechiniscus ramazzottii lineatus Maucci, 1973-74 see Pseudechiniscus ramazzottii facettalis
*Pseudechiniscus shilinensis Yang, 2002
Pseudechiniscus sinensis Rahm, 1937

Testechiniscus spinuloides (Murray, 1907) see Testechiniscus spitsbergensis

Testechiniscus spitsbergensis (Scourfield, 1897) [Echiniscus clavisetosus Mihelčič, 1958 in Dastych 1973a; Echiniscus
marinellae Bartoš, 1935 in Petersen 1951, Maucci 1985; Echiniscus melanophthalmus Bartoš, 1936 in Petersen
1951, Maucci 1985; Echiniscus menzeli Heinis, 1917 in Cuénot 1932, Petersen 1951, Maucci 1985; Echiniscus
rosaliae Mihelčič, 1951 in Ramazzotti & Maucci 1983, Maucci 1985; Echiniscus spinuloides Murray, 1907 in
Cuénot 1932, Petersen 1951, Maucci 1985]

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p. 27
*THERMOZODIA Ramazzotti & Maucci, 1983

*Milnesium almatyense Tumanov, 2006

*Milnesium antarcticum Tumanov, 2006
*Milnesium asiaticum Tumanov, 2006
Milnesium eurystomum Maucci, 1991
*Milnesium longiungue Tumanov, 2006
*Milnesium reductum Tumanov, 2006

p. 28
Amphibolus nebulosus Dastych, 1983 {in the description of the species (Dastych 1983), due to printing error the firstly
used name was nebolosus but in the rest of the text the name nebulosus was used}
Amphibolus smreczynskii (Węglarska, 1970)

Eohypsibius nadjae Kristensen, 1982 {egg described in Bertolani et al. 1996, Guidetti & Bertolani 2001a}

Calohypsibius ornatus (Richters, 1900) [Hypsibius (Calohypsibius) armatus Bartoš, 1938; Hypsibius (Calohypsibius)
intermedius Mihelčič, 1939] {see Pilato et al. 1989}
*Calohypsibius maliki Michalczyk & Kaczmarek, 2005
Calohypsibius placophorus (da Cunha, 1943)

Haplohexapodibius seductor Pilato & Beasley, 1987 [Hexapodibius beasleyi Maucci, 1988 in Kaczmarek et al. 2006]

*Haplomacrobiotus hermosillensis May, 1948 <Amended by Pilato 1973c>

*Haplomacrobiotus utahensis Pilato & Beasley, 2005

Hexapodibius Pilato, 1969

<Amended by Schuster et al. 1980> {Maucci (1981) suggested the unification of both genera Hexapodibius and
Parhexapodibius and gave a new diagnosis for the genus Hexapodibius}
Hexapodibius beasleyi Maucci, 1988 see Haplohexapodibius seductor
Hexapodibius boothi Dastych & McInnes, 1994

Parhexapodibius Pilato, 1969

<Amended by Schuster et al. 1980> {Ramazzotti & Maucci (1983) did not consider valid Parhexapodibius and
inserted it in Hexapodibius; Nelson & McInnes (2002) consider these two genera as valid}

p. 29
Acutuncus Pilato & Binda, 1997
Acutuncus antarcticus (Richters, 1904) <Amended by Dastych 1991> [Hypsibius simoizumii Sudzuki, 1964 in Dastych
1991]

*Borealibius Pilato, Guidetti, Rebecchi, Lisi, Hansen & Bertolani, 2006

Borealibius zetlandicus (Murray, 1907) <Transferred from Hypsibius to Borealibius by Pilato et al. 2006>

Doryphoribius flavus (Iharos, 1966) [Doryphoribius citrinus (Maucci, 1973) in Schuster & Toftner 1982]
Doryphoribius pilatoi R. Bertolani, 1984
*Doryphoribius qinlingense Li, Su & Yu, 2004
Doryphoribius vietnamensis (Iharos, 1969) <Transferred from Isohypsibius to Doryphoribius by Beasley et al. 2006>
Doryphoribius zyxiglobus (Horning, Schuster & Grigarick, 1978)

Halobiotus stenostomus (Richters, 1908) [Macrobiotus appelloefi Richters, 1908 in Marcus 1936; {Hypsibius (Isohyps-
ibius) geddesi Hallas, 1971 synonymus of H. appelloefi by Petelina & Tchesunov (1983) who consider valid this last
species}]

p. 30
*Hypsibius choucoutiensis Rahm, 1937
*Hypsibius heardensis Miller, McInnes & Bergstrom, 2005
Hypsibius iharosi Bartoš, 1941 see Hypsibius montanus

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*Hypsibius iskandarovi Tumanov, 1997

Hypsibius klebelsbergi Mihelčič, 1959 <Amended by Dastych 1993, Dastych et al. 2003 >
*Hypsibius kunmingensis Yang, 2002
Hypsibius montanus Iharos, 1940 [Hypsibius iharosi Bartoš, 1941 syn. nov.] {combination Hypsibius montanus was
used first by Iharos (1940) and not by Mihelčič (1938b) as declared by Bartoš (1941) when he renamed this species
as Hypsibius iharosi}
Hypsibius septulatus Pilato, Binda, Napolitano & Moncada, 2004
*Hypsibius seychellensis Pilato, Binda & Lisi, 2006
Hypsibius simoizumii Sudzuki, 1964 see Acutuncus antarcticus
*Hypsibius thaleri Dastych, 2004
Hypsibius zetlandicus (Murray, 1907) see Borealibius zetlandicus

*Isohypsibius altai Kaczmarek & Michalczyk, 2006

Isohypsibius annulatus minor (Ramazzotti, 1945)

p. 31
Isohypsibius arcuatus (Bartoš, 1934)
Isohypsibius baicalensis (Ramazzotti, 1966)
Isohypsibius baldii (Ramazzotti, 1945)
*Isohypsibius baldiioides Tumanov, 2003
Isohypsibius basalovoi (Durante & Maucci, 1973)
Isohypsibius belliformis (Mihelčič, 1971)
Isohypsibius bellus (Mihelčič, 1971)
*Isohypsibius borkini Tumanov, 2003
*Isohypsibius brevitubulatus Rho, Chang & Kim, 1997 (in Rho et al. 1997)
Isohypsibius brulloi Pilato & Pennisi, 1976
Isohypsibius bulbifer (Mihelčič, 1957)
Isohypsibius cameruni (Iharos, 1969)
*Isohypsibius changbaiensis Yang, 1999
Isohypsibius costatus (Mihelčič, 1971)
Isohypsibius cyrilli (Mihelčič, 1951)
Isohypsibius deflexus (Mihelčič, 1960)
Isohypsibius duranteae (Maucci, 1978)
Isohypsibius effusus (Mihelčič, 1971)
Isohypsibius franzi (Mihelčič, 1951)
Isohypsibius fuscus (Mihelčič, 1971/72)
Isohypsibius gibbus (Marcus, 1928) see Isohypsibius verrucosus
Isohypsibius glaber (Durante Pasa & Maucci, 1979)
Isohypsibius gracilis (Iharos, 1966)
Isohypsibius gyulai (Mihelčič, 1971)
Isohypsibius hadzii (Mihelčič, 1938)
Isohypsibius hypostomoides (Mihelčič, 1971)

p. 32
Isohypsibius itoi Tsurusaki, 1980 see Thulinius itoi
*Isohypsibius jingshanensis Yang, 2003
*Isohypsibius jinhouensis Yang, 2007
*Isohypsibius kotovae Tumanov, 2003
*Isohypsibius ladogensis Tumanov, 2003
Isohypsibius landalti (Iharos, 1966) nomen nudum
Isohypsibius leithaicus (Iharos, 1966)
*Isohypsibius liae Li & Wang, 2006
Isohypsibius lineatus (Mihelčič, 1969)
Isohypsibius macrodactylus (Maucci, 1978) [Isohypsibius zierhofferi Dastych, 1980]
Isohypsibius marii R. Bertolani, 1982
Isohypsibius monoicus R. Bertolani, 1982
Isohypsibius monstruosus Maucci, 1991
Isohypsibius myrops (du Bois-Reymond Marcus, 1944)
Isohypsibius neoundulatus (Durante Pasa & Maucci, 1975)

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Isohypsibius nipponicus Sudzuki, 1975

*Isohypsibius panovi Tumanov, 2005
Isohypsibius papillifer bulbosus (Marcus, 1928)
Isohypsibius pauper (Mihelčič, 1971)
Isohypsibius pilatoi (Durante Pasa & Maucci, 1979)
Isohypsibius prosostomus cambrensis (Morgan, 1976)
Isohypsibius pseudoundulatus (da Cunha & do Nascimento Ribeiro, 1964)
Isohypsibius pulcher (Mihelčič, 1971/72)
*Isohypsibius pushkini Tumanov, 2003
*Isohypsibius qinlingensis Li, Wang & Yu, 2005
*Isohypsibius rahmi Li & Wang, 2006

p. 33
Isohypsibius sabellai Pilato, Binda, Napolitano & Moncada, 2004
Isohypsibius saltursus Schuster, Toftner & Grigarick, 1978
Isohypsibius sattleri (Richters, 1902) {corrected according to Marcus 1936} [Hypsibius bakonyiensis Iharos, 1964 in
Dastych 1990] <Amended by Pilato 1973b; Dastych 1990>
Isohypsibius schaudinni (Richters, 1909)
Isohypsibius sculptus (Ramazzotti, 1962) <Amended by Dastych 1997>
Isohypsibius sellnicki (Mihelčič, 1962)
Isohypsibius sismicus (Maucci, 1978)
Isohypsibius solidus (Mihelčič, 1971)
*Isohypsibius taibaiensis Li & Wang, 2005
Isohypsibius tetradactyloides (Richters, 1907)
Isohypsibius torulosus (Mihelčič, 1959)
Isohypsibius tuberculatus (Plate, 1888)
Isohypsibius tuberculoides (Mihelčič, 1951) {in Mihelčič 1951, both names tuberculuides and tuberculoides were used
but in a later description (Mihelčič 1953), only tuberculoides was used}
Isohypsibius vejdovskyi (Bartoš, 1939)
Isohypsibius verrucosus (Della Valle, 1915) [Isohypsibius gibbus (Marcus, 1928) syn. nov.] {Marcus (1928) transferred
Macrobiotus tuberculatus var. verrucosus Della Valle, 1915 to the genus Hypsibius and renamed it to Hypsibius
tuberculatus f. gibbus to avoid confusion with Calohypsibius verrucosus (Richters, 1900), which was also in the
same genus Hypsibius at that time. Therefore, the valid name of this species is Isohypsibius verrucosus (Della Valle,
1915)}
Isohypsibius vietnamensis (Iharos, 1969) see Doryphoribius vietnamensis
*Isohypsibius yunnanensis Yang, 2002
Isohypsibius zierhofferi Dastych, 1980 see Isohypsibius macrodactylus

Pseudobiotus longiunguis (Iharos, 1968) {formerly Echinursellus longiunguis according to Kristensen 1987}

p. 34
Ramazzottius affinis Bertolani, Guidetti & Rebecchi, 1994
*Ramazzottius bunikowskae Kaczmarek, Michalczyk & Diduszko, 2006
*Ramazzottius nivalis Dastych, 2006
Ramazzottius szeptycki (Dastych, 1980)

Thulinius R. Bertolani, 2003

[Thulinia R. Bertolani, 1982 in Bertolani 2003] <Amended by Bertolani et al. 1999>
Thulinius itoi (Tsurusaki, 1980) <Transferred from Isohypsibius to Thulinius by Kaczmarek & Michalczyk 2006a>
Thulinius ruffoi (R. Bertolani, 1982) <Amended by Bertolani et al. 1999>

Diphascon (Diphascon) Plate, 1888 {corrected according to Marcus 1936}

<Amended by Pilato 1987>
Diphascon (Diphascon) aculeatum (Maucci, 1951-1952)
Diphascon (Diphascon) australianum Pilato & Binda, 1998
Diphascon (Diphascon) birklehofi Rolf Schuster, 1999
Diphascon (Diphascon) bisbullatum (Iharos, 1964) {to be confirmed in this subgenus, see Pilato 1987}

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p. 35
Diphascon (Diphascon) brevipes (Marcus, 1936)
*Diphascon (Diphascon) dolomiticum Pilato & Bertolani, 2005
Diphascon (Diphascon) elongatum (Mihelčič, 1959) {to be confirmed in this subgenus, see Pilato 1987}
Diphascon (Diphascon) gerdae (Mihelčič, 1951) {to be confirmed in this subgenus, see Pilato 1987}
Diphascon (Diphascon) halapiense (Iharos, 1964) {to be confirmed in this subgenus, see Pilato 1987}
Diphascon (Diphascon) humicus Bertolani, Guidetti & Rebecchi, 1995
*Diphascon (Diphascon) hydrophilum Pilato, Binda, Bertolani & Lisi, 2005
Diphascon (Diphascon) iltisi (Schuster & Grigarick, 1965)
Diphascon (Diphascon) langhovdense (Sudzuki, 1964) {corrected according to Dastych 2002/2003}
Diphascon (Diphascon) latipes (Mihelčič, 1955) {to be confirmed in this subgenus, see Pilato 1987}
Diphascon (Diphascon) mirabilis Dastych, 1984
Diphascon (Diphascon) mitrense Pilato, Binda & Qualtieri, 1999
*Diphascon (Diphascon) nelsonae Pilato, Binda, Bertolani & Lisi, 2005
Diphascon (Diphascon) nobilei (Binda, 1969)
Diphascon (Diphascon) nodulosum (Ramazzotti, 1957)
Diphascon (Diphascon) nonbullatum (Mihelčič, 1951) {to be confirmed in this subgenus, see Pilato 1987}
Diphascon (Diphascon) oculatum oculatum Murray, 1906
Diphascon (Diphascon) oculatum alpinum (Mihelčič, 1964)
Diphascon (Diphascon) oculatum canadense Murray, 1910 [Hypsibius vancouverensis Thulin, 1911] {Diphascon ocula-
tum vancouverense Thulin, 1911 was firstly described as Diphascon canadense Murray, 1910 but renamed by Thulin
(1911) to Hypsibius vancouverensis through the homonymy with Isohypsbius canadensis (Murray, 1910) as these
two species were in the same genus Hypsibius according to Thulin (1911). Marcus (1936) considered it to be only a
form of Hypsibius (Diphascon) oculatus. Therefore, the current valid name of this taxon is Diphascon (Diphascon)
oculatum canadense Murray, 1910}
Diphascon (Diphascon) oculatum vancouverense Thulin, 1911 see Diphascon (Diphascon) oculatum canadense
Diphascon (Diphascon) ongulense (Morikawa, 1962)
Diphascon (Diphascon) pingue pingue (Marcus, 1936) <Amended by Pilato & Binda 1997/98>
*Diphascon (Diphascon) platyungue Pilato, Binda, Bertolani & Lisi, 2005
Diphascon (Diphascon) recamieri Richters, 1911
Diphascon (Diphascon) rugocaudatum (Rodriguez Roda, 1952)
Diphascon (Diphascon) rugosum (Bartoš, 1935)

p. 36
*Diphascon (Diphascon) serratum Pilato, Binda, Bertolani & Lisi, 2005
Diphascon (Diphascon) stappersi Richters, 1911 {to be confirmed in this subgenus, see Pilato 1987}
Diphascon (Diphascon) trachydorsatum (Bartoš, 1937)

Diphascon (Adropion) arduifrons Thulin, 1928

Diphascon (Adropion) clavatum (Bartoš, 1935)
Diphascon (Adropion) onorei Pilato, Binda, Napolitano & Moncada, 2002
Diphascon (Adropion) scoticum scoticum Murray, 1905 {belonging to the genus Fujiscon according to Degma &
Opravilová 2001} [Diphascon crozetense Richters, 1907]
Diphascon (Adropion) scoticum ommatophorum (Thulin, 1911) {corrected according to Marcus 1936}
*Diphascon (Adropion) scoticum qinlingensis Li & Liu, 2005

Diphascon coniferens (Bartoš, 1960) {subgenus to be defined}

Diphascon mariae (Mihelčič, 1951) {subgenus to be defined}
Diphascon punctatum (Iharos, 1962) {subgenus to be defined}

Hebesuncus conjungens (Thulin, 1911) <Amended by Dastych & Thaler 2002>

p. 37
Itaquasconinae Rudescu, 1964 {for a discussion on the authorship of the taxon see Dastych 1992}

Astatumen bartosi (Węglarska, 1959)

Astatumen trinacriae (Arcidiacono, 1962) [Itaquascon ramazzottii Iharos, 1966 in Pilato 1969c]

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Itaquascon de Barros, 1939

<Amended by Pilato 1997>
Itaquascon cambewarrense Pilato, Binda & Claxton, 2002
Itaquascon mongolicus Kaczmarek, Michalczyk & Węglarska, 2002
Itaquascon pawlowskii Węglarska, 1973
Itaquascon placophorum Maucci, 1973 {in the genus Astatumen in Pilato 1997 but confirmed in the genus Itaquascon by
Guidetti et al. 1999}
Itaquascon umbellinae de Barros, 1939

Platicrista itaquasconoide (Durante Pasa & Maucci, 1975) {to be confirmed in this genus, see Pilato 1987}
*Platicrista ramsayi Marley, 2006

p. 38
Macrobiotinae Guidetti, Rebecchi & Bertolani, 2000 {subfamily not longer valid, see Guidetti et al. 2005}

Biserovus Guidetti & Pilato, 2003

*Calcarobiotus (Calcarobiotus) longinoi Kaczmarek, Michalczyk & Guidetti, 2006

Calcarobiotus (Discrepunguis) Guidetti & Bertolani, 2001

Calcarobiotus (Discrepunguis) polygonatus (Binda & Guglielmino, 1991) [Macrobiotus eugranulatus Maucci, 1993 in
Pilato et al. 2004]

Insuetifurca Guidetti & Pilato, 2003

*Insuetifurca austronipponica Abe, 2005

Macrobiotus acontistus de Barros, 1942 see Minibiotus acontistus

*Macrobiotus alekseevi Tumanov, 2005
Macrobiotus allani Murray, 1913 see Minibiotus allani
*Macrobiotus alvaroi Pilato & Kaczmarek, 2007
*Macrobiotus aradasi Binda, Pilato & Lisi, 2005
Macrobiotus ariekammensis Węglarska, 1965 [Macrobiotus adelges Dastych, 1977 in Dastych 1985]

p. 39
*Macrobiotus barabanovi Tumanov, 2005
Macrobiotus beotiae Durante Pasa & Maucci, 1979
Macrobiotus biserovi Bertolani, Guidi & Rebecchi, 1996
Macrobiotus brevipes Mihelčič, 1971/72
Macrobiotus cargatensis Pilato, Binda & Lisi, 2002 see Macrobiotus corgatensis
*Macrobiotus contii Pilato & Lisi, 2006
Macrobiotus coronatus de Barros, 1942 <Amended by Pilato et al. 2000>
Macrobiotus crassidens Murray, 1907 see Minibiotus crassidens
Macrobiotus crenatus Maucci, 1991
Macrobiotus danielae Pilato, Binda, Napolitano & Moncada, 2001
*Macrobiotus danielisae Pilato, Binda & Lisi, 2006
*Macrobiotus danilovi Tumanov, 2007
*Macrobiotus divergens Binda, Pilato & Lisi, 2005
Macrobiotus dubius Murray, 1907 species inquirenda {this species was omited in Ramazzotti & Maucci 1983 as it was
impugned in Ramazzotti 1972}
Macrobiotus evelinae de Barros, 1938
Macrobiotus furciger Murray, 1907 [Macrobiotus ehrenbergi Heinis, 1921] {formerly Macrobiotus furcatus Murray,
1906 but later renamed by Murray (1908) through the homonymy with Macrobiotus furcatus Ehrenberg, 1859}
*Macrobiotus garynahi Kaczmarek, Michalczyk & Diduszko, 2005
Macrobiotus gemmatus Bartoš, 1963 {it should be better studied for a possible transfer to Minibiotus}
Macrobiotus granatai Pardi, 1941 see Minibiotus granatai
*Macrobiotus grandis Richters, 1911 <Amended by Maucci & Pilato 1974>
*Macrobiotus hapukuensis Pilato, Binda & Lisi, 2006
Macrobiotus hibiscus de Barros, 1942
*Macrobiotus hufelandi hufelandi C.A.S. Schultze, 1833 <Amended by Bertolani & Rebecchi 1993>

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*Macrobiotus huziori Michalczyk & Kaczmarek, 2006

Macrobiotus hyperonyx Maucci, 1983

p. 40
*Macrobiotus insularis Pilato, 2006
Macrobiotus islandicus islandicus Richters, 1904 [Macrobiotus ruffoi Maucci, 1973]
*Macrobiotus kirghizicus Tumanov, 2005
Macrobiotus kurasi Dastych, 1981
Macrobiotus lazzaroi Maucci, 1986 {formerly Macrobiotus spallanzanii Maucci, 1973 but later renamed by Maucci
(1986) through the homonymy with Macrobiotus spallanzanii Della Valle, 1915}
Macrobiotus longipes Mihelčič, 1971/72
Macrobiotus madegassus Maucci, 1993
*Macrobiotus magdalenae Michalczyk & Kaczmarek, 2006
*Macrobiotus mandalaae Pilato, 1974 <Amended by Pilato 1984, 2006>
Macrobiotus mongolicus Maucci, 1988
Macrobiotus montanus Murray, 1910 [Macrobiotus morulatus Bartoš, 1936]
Macrobiotus morulatus Bartoš, 1936 see Macrobiotus montanus
Macrobiotus norvegicus Mihelčič, 1971/72
Macrobiotus occidentalis occidentalis Murray, 1910 {it should be better studied for a possible transfer to Minibiotus}
*Macrobiotus ocotensis Pilato, 2006
Macrobiotus ovostriatus Pilato & Patanè, 1998
Macrobiotus peterseni Maucci, 1991
Macrobiotus picardi Mihelčič, 1960 species inquirenda {this species was omited in Ramazzotti & Maucci 1983 as it was
impugned in Ramazzotti 1972}

p. 41
Macrobiotus potockii Węglarska, 1968
Macrobiotus primitivae de Barros, 1942
Macrobiotus psephus du Bois-Reymond Marcus, 1944
*Macrobiotus pseudocoronatus Pilato, Binda & Lisi, 2006
Macrobiotus pseudofurcatus Pilato, 1972
*Macrobiotus ramoli Dastych, 2005
Macrobiotus richtersi Murray, 1911 [Macrobiotus schultzei Greeff, 1866] {corrected according to Cuénot 1932 and Mar-
cus 1936}
*Macrobiotus rigidus Pilato & Lisi, 2006
*Macrobiotus semmelweisi Pilato, Binda & Lisi, 2006
Macrobiotus serratus Bertolani, Guidi & Rebecchi, 1996
*Macrobiotus shennongensis Yang, 1999
*Macrobiotus siamensis Tumanov, 2006
*Macrobiotus sicheli Binda, Pilato & Lisi, 2005
*Macrobiotus sklodowskae Michalczyk, Kaczmarek & Węglarska, 2006
*Macrobiotus spectabilis Thulin, 1928 <Amended by Dastych 1973b, Maucci & Pilato 1974> {possible synonym of
Macrobiotus grandis Richters, 1911 in Dastych, 1973b}
Macrobiotus stellaris du Bois-Reymond Marcus, 1944
Macrobiotus striatus Mihelčič, 1949
*Macrobiotus tenuiformis Tumanov, 2007
Macrobiotus topali Iharos, 1969 {it should be better studied for a possible transfer to Minibiotus}
Macrobiotus tridigitus R.O. Schuster, 1983 see Schusterius tridigitus
*Macrobiotus voronkovi Tumanov, 2007
*Macrobiotus yunshanensis Yang, 2002

p. 42
Minibiotus R.O. Schuster, 1980 (in Schuster et al. 1980)
<Amended by Claxton 1998>
Minibiotus acontistus (de Barros, 1942) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
Minibiotus aculeatus (Murray, 1910) [Macrobiotus intermedius subjulietae Horning, Schuster & Grigarick, 1978 in
Claxton, 1998]
Minibiotus africanus Binda & Pilato, 1995 {egg described in Binda et al. 2001}
Minibiotus allani (Murray, 1913) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>

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Minibiotus constellatus Michalczyk & Kaczmarek, 2003

*Minibiotus continuus Pilato & Lisi, 2006
Minibiotus crassidens (Murray, 1907) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
Minibiotus diphasconides (Iharos, 1969) {for comments on the species name and status see Guidetti & Pilato 2003}
*Minibiotus eichhorni Michalczyk & Kaczmarek, 2004
Minibiotus furcatus (Ehrenberg, 1859) [nec Macrobiotus furcatus Murray, 1906] <Amended by Binda & Pilato 1992;
Claxton 1998>
Minibiotus granatai (Pardi, 1941) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
*Minibiotus gumersindoi Guil & Guidetti, 2005
Minibiotus intermedius (Plate, 1888) <Amended by Claxton 1998>
Minibiotus julietae (de Barros, 1942) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
Minibiotus lazzaroi (Maucci, 1986) see Macrobiotus lazzaroi {erroneously transferred to Minibiotus by Guidetti &
Bertolani 2005}
Minibiotus marcusi (de Barros, 1942) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
Minibiotus pseudofurcatus (Pilato, 1972) see Macrobiotus pseudofurcatus {erroneously transferred to Minibiotus by
Guidetti & Bertolani 2005}
Minibiotus pustulatus (Ramazzotti, 1959) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
Minibiotus striatus (Mihelčič, 1949) see Macrobiotus striatus {erroneously transferred to Minibiotus by Guidetti &
Bertolani 2005}
Minibiotus subintermedius (Ramazzotti, 1962) <Transferred from Macrobiotus to Minibiotus by Guidetti et al. 2007>
Minibiotus subjulietae Horning, Schuster & Grigarick, 1978 see Minibiotus aculeatus
*Minibiotus weglarskae Michalczyk, Kaczmarek & Claxton, 2005

Minilentus Guidetti & Pilato, 2003

Pseudodiphascon inflexum (Arcidiacono, 1964) species inquirenda [in Guidetti & Pilato 2003]

p. 43
*Schusterius Kaczmarek & Michalczyk, 2006
Schusterius tridigitus (R.O. Schuster, 1983) <Transferred from Macrobiotus to Schusterius by Kaczmarek & Michalc-
zyk 2006b>

Murrayidae Guidetti, Gandolfi, Rossi & Bertolani, 2005 {formerly Murrayinae Guidetti, Rebecchi & Bertolani,
2000}

Dactylobiotus R.O. Schuster, 1980 (in Schuster et al. 1980)

*Dactylobiotus aquatilis Yang, 1999
Dactylobiotus grandipes (Schuster, Toftner & Grigarick, 1978)
Dactylobiotus haplonyx Maucci, 1981
*Dactylobiotus henanensis Yang, 2002
*Dactylobiotus octavi Guidetti, Altiero & Hansen, 2006
Dactylobiotus parthenogeneticus R. Bertolani, 1982
Dactylobiotus selenicus R. Bertolani, 1982

p.44
Apodibius nuntius Binda, 1984 [Apodibius serventyi Morgan & Nicholls, 1986 in Van Rompu et al. 1995]
Apodibius serventyi Morgan & Nicholls, 1986 see Apodibius nuntius

References

Guidetti, R. & Bertolani, R. (2005) Tardigrade taxonomy: an updated checklist of the taxa and a list of characters for
their identification. Zootaxa, 845, 1–46.
International Code of Zoological Nomenclature. Fourth Edition (1999) London: The International Trust for Zoological
nomenclature, xxix+306 pp.
McInnes, S.J. (1994) Zoogeographic distribution of terrestrial/freshwater tardigrades from current literature. Journal of
Natural History, 28, 257–352.
Ramazzotti, G. & Maucci, W. (1983) Il Phylum Tardigrada. Terza edizione riveduta e corretta. Memorie dell’Isituto
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