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Structural Features
Concept of Ecosystem
Biotic structures
An ecosystem is an integrated unit consisting of
(a) Producers photoautotrophs : mainly green plants
interactions between microorganisms, plants, animals
- generate energy and food from water,
whose survival depends on the maintenance and carbondioxide in air and importantly sunlight
regulation
l ti off their
th i biotic
bi ti and
d abiotic
bi ti structure
t t and
d function.
f ti
- Chemoautotrophs : some microorganisms and
Ecosystem large variations in size, structure, deep sea organisms
composition,etc. - generate energy from inorganic reduction rxns
All ecosystems have certain basic structural and functional - Chemo/phototrophs : Some microorganisms
which use solar energy as well as chemical
features which are common energy
Ecosystem
Ecosystem
Structural Features
Biotic structures Structural Features
(b) Consumers (heterotrophs) Organisms which get their
organic food by feeding on other organisms Abiotic structures
Herbivores (primary consumers)
(a) Physical factors sunlight, temperature, rainfall, wind,
Carnivores (secondary and tertiary consumers)
latitude/longitude soil,
latitude/longitude, soil water,
water
Omnivores (feed both on plants and animals)
Detritivores (saprotrophs) (b) Chemical factors major and minor essential nutrients like
Decomposers (microbial systems) carbon, nitrogen, phosphorous, potassium, hydrogen, oxygen,
In all ecosystems, the biotic structure prevails but the
dominance of any component depends on the ecosystems sulphur
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Ecosystem Ecosystem
Functional Features Functional Features
(a) Food chain, food web and trophic structure (a) Food chain, food web and trophic structure
(e) Ecosystem development and regulation ecosystem and their interaction of energy transfer along with
population size - Trophic structure
Ecosystem
Level 4 Ecosystem
Functional Features
(a) Food chain -a (a) Food chain - all organisms, living or dead, are potential food
sequence of eating Level 3
for some other organism
and being eaten
Ex. Desert ecosystem The detritus food chain also exist which constitutes detrivores
Level 2 and decomposers
g
Each organism in the
ecosystem is assigned
The surface food chain derives its energy basically from plant
a feeding level or
trophic level Level 1 energy while the detritus food chain primarily obtains energy
depending on its from plant biomass, secondarily from microbial biomass and
nutritional status
tertiarily from carnivores.
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Ecosystem
Functional Features
(a) Food webs shows pattern of energy or nutrient flow through Ecosystem
out the community
(a) Food web network of food chains where different types of
organisms are connected at different trophic levels, so that
there are a number of options of eating and being eaten at
each trophic level.
Biomagnification
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Ecological pyramids Pyramid of numbers
Ecological pyramids are graphical representations of the number Represents the number of
individual organisms at
of individuals in different nutritional levels. For example, the plant- each trophic level
insect-bird-hawk food chain can be represented as an ecological The pyramid may be
pyramid upright or inverted based
on the ecosystem
Plants absorb energy from the sun, the insects eat the plants, the
Pond, grassland, forest
birds eat the insects,, and the hawks eat the birds. Hence,, the upright
energy of the sun has been transferred from the sun to the tissues Parasitic ecosystem -
of the hawk. Since the number of individuals in each level usually inverted
decreases, the resulting diagram looks like a pyramid.
microbes Hyper-parasites
Pyramid of numbers Lice and bugs parasites
birds herbivores
Pyramid of Biomass trees producers
Pyramid of energy
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Energy Flow in an Ecosystem
Single channel energy flow model
Flow of energy in an ecosystem takes place through the food chain
and it is this energy flow which keeps the ecosystem functional
Unused energy Unused energy
The important feature of this flow is that it is Unidirectional or one sunlight
way flow Energy NA
Energy NA
Universal energy flow model Unused energy
Input
gy
energy GPP NPP Output
Energy storage
energy
Carnivores
producers herbivores
Living Biomass Model depicts the gradual decline in energy level due to loss of energy
at each successive trophic level in a grazing food chain
respiration
Primary productivity is the rate at which radiant energy is converted In nature the surface (grazing) food chain and detritus food chain
into organic substances by photosynthesis by primary producers. operate in the same ecosystem
Plants must use organic molecules to fuel their own cellular
The volume of primary production passed to detritus food chain varies
respiration, about 55%
with ecosystem. Ex. In forest ecosystem huge quantity of
55% of gross primary productivity is available to heterotrophs; this
biomass produced cannot be all consumed by herbivores and
is net primary productivity
most of it p
passes into detritus food chain;; In marine ecosystem
y
Secondary productivity the energy stored at consumer level for there is very little primary production passed to detritus food chain
use by the next trophic level is defined as secondary production. Energy flow models involving both food chains Double channel or
The secondary production is the amount of organic matter stored Y-shaped energy food chain
by the herbivores or carnivores (in excess of respiratory and un-
assimilated energy loss).
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Trophic cascade: effect of one trophic level flow up or down to Energy flow in an ecosystem is a consequence of two
more than one adjacent trophic level. Ex: a predator (lion) not only fundamental laws of thermodynamics
affects its prey population (impala) but, by its eating of the prey, First law of thermodynamics - energy can neither be created
also the preys prey (grass) nor destroyed; it can only be changed from one form of energy to
another
Top-Down effects: predators control lower trophic level Second law of thermodynamics - when energy is transformed
populations Ex: Despite the presence of herbivores, not all plant from one form to another, there is always some loss of energy
life is consumed. Predators keep herbivore population to a level from the system, usually as low grade heat
that the herbivores cannot extinct the plants
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CARBON CYCLE:
Sedimentary cycles: Lithosphere is reservoir cybernetically Principle building block of all kinds of molecules which make up
living organisms. Most significant element for existence and
less controlled mobility of materials in the reservoir is survival of life on earth.
relatively less and cycles are slow. Carbon is present in air (as CO2), in water (as dissolved CO2,
H2CO3, HCO3 and CO3), in rocks (as CO3), in fossil fuels (as
- Less perfect because the elements get locked in reservoir and petroleum and coal), in all life forms (as proteins, fats and
carbohydrates)
go out of circulation for long periods.
83% of C is present as inorganic minerals in the rocks of earth and
- Elements such as P,
P S,
S K and Ca have sedimentary cycles.
cycles rate at which it is released is extremely slow.
slow
Most predominant movement of C in the carbon cycle is through
- Human beings can influence the nutrient cycling (a mighty the biotic processes of photosynthesis and respiration.
geological agent) can make material movement faster and Average concentration of CO2 in atmosphere is 320 ppm. The
material cycles imperfect. oceanic reservoir is 50 times larger. About 3 x 1010 metric
tonnes of C is converted to 12x 1010 metric tonnes of sugar
- Human influence is maximum felt on the smallest pool of the annually.
NITROGEN CYCLE
Nitrogen is the component of proteins, nucleic acids and ATP which
are essential structural and functional components of living
systems.
Nitrogen cycle is a gaseous cycle includes Nitrogen fixation,
Nitrification, Nitrogen assimilation, ammonification and
denitrification in cycling nitrogen in nature from Environment to
organisms and from organisms back to environment.
Nitrogen Fixation : Elemental Nitrogen of atmosphere is
converted into Nitrogenous compound - It takes place by two
methods.
(1) Physico-Chemical Nitrogen Fixation:
Nitrogen from the atmosphere is fixed in the form of oxides as a
result of lightning and volcanic eruptions. 10% Nitrogen is fixed
in this way.
(2) Biological Nitrogen Fixation: - By free living bacteria like
Azotobacter, Clostridium, Blue green algae like Nostoc,
Anabaena,Cylindrospermum.
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By symbiotic bacteria like Rhizobium found in the root nodules of
leguminous plants. - Incorporation of Ammonia into amino acids which in turn from
proteins. Animals take proteins from the plants which supply
N2 + 3H2 2NH3 amino acids necessary for the metabolism of animals.
Nitrification : Conversion of ammonia into nitrates is called Ammonification: Conversion of urea and uric acid excreted by
nitrification. Ammonia is first converted into nitrites by bacteria animals and proteins from dead plants and animals into
Nitrosomonas & Nitrite is further converted into nitrates by ammonia by ammonifying or putrefying bacteria like Clostridium,
Nitrobacter. Proteus, Pseudomonas, Bacillus etc.
Nitrosomonas Denitrification : Reduction of Nitrate ions in the soil into nitrogen
2NH3 + O2 2 HNO2 + 2H2O + Energy gas by denitrifying bacteria such as Pseudomonas, Thiobacillus
denitrificans. This nitrogen gas is once again released into the
Nitrobacter atmosphere.
HNO2 + O2 HNO3 + Energy Occurs in water logged soils where anaerobic conditions exists.
Nitrogen Assimilation: Reduction of nitrates into ammonia in the NO3 NO2 N2O N2
plant body.
PHOSPHOROUS CYCLE
Phosphorous cycle is a sedimentary cycle with major reserves of
phosphorous in the sediments
The reservoir of phosphorous lies in the rocks, fossils, etc.
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NUTRIENT CYCLES IN TROPICAL & TEMPERATE REGIONS
- Root mat root for recovering nutrients from the surface
Temperate systems have greater proportion of nutrient in the litter
soil or sediments while tropical systems have in the biomass.
- Mycorrhizae acting as nutrient traps
Tropical Temperate - Algae & lichens on plant fixing nitrogen and scavenging
Vegetation Above Vegetation Above nitrogen from rain water.
Ground Ground - Disturbance to the system/biotic community destroys the
Carbon 7.5% -- 50% -- mechanisms for cycling of nutrient within the biotic pool
Nitrogen 58% 44% 9% 6% and results in their loss to abiotic pool.
Nutrient cycles tends to be physical in temperate regions while - Nutrients are ultimately lost from the abiotic pool due to
it is biological in tropical regions. prevailing high temperature and rainfall.
Tropical soils are nutrient poor and nutrients mostly cycle - High temp. results in rapid mineralization of nutrients.
within the organic structure of the system. Rich species - High rain fall leads to leaching and runoff loss of nutrients
diversity and symbiotic associations between autotrophs and
heterotrophs including the following prevents loss of nutrients
from the biotic pool.
ECOSYSTEM STABILITY
ATTRIBUTES RELATED TO NUTRIENT CYCLING IN
ECOSYSTEM:
Stability is the ability to resist change when perturbed
two types
Cycling index : Mineral cycles are open initially but become
increasingly closed. - Resistance Stability ability to withstand external
Nutrient exchange rapid initially becomes slow at later stage. disturbance without getting perturbed.
Turnover time and storage of essential elements increases - Resilience Stability ability of the perturbed ecosystem by
during succession and reach a steady state.
external disturbances to return to original state once the
Nutrient retention & conservation poor initially but increases disturbance is removed.
and becomes pronounced later.
Benign environment supports development of systems with
resistance stability while variable environmental conditions
favour development of resilience stability.
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Servo mechanisms and homeostatic mechanism of control:
Death or collapse
Servo Mechanism - control system is external to the system
being controlled.
Homeostatic mechanism - control system is internal to the + ve feedback
system being controlled, two types - centralized and
distributed/diffused control.
Homeostatic
Control mechanisms can be either feed back and feed forward -ve
ve feedback -ve
ve feedback
plateau
l
types.
Feed backs may be positive or negative.
+ ve feedback
Ecosystems are known for redundancy believed to have
both servo and homeostatic mechanisms (diffused type) of
control.
Death or collapse
Stress
(-) (0) (+) conditions
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ATTRIBUTES OF SUCCESSION:
TYPES OF SUCCESSION:
A multitude of attributes can be used to describe changes associated
1. Primary succession & secondary succession with the succession.
2. Autotrophic & Heterotrophic succession Degree and rate of change and time required to reach steady state.
Autotrophic succession occurs in a medium rich in organic - May vary from attribute to attribute
substances and starts with P<R situation - May be influenced by climate
Heterotrophic succession begins in a medium rich in organic matter ATTRIBUTES RELATED TO ENERGY FLOW THROUGH
and starts with P>R situation. ECOSYSTEM (ENERGETICS)
Succession tends to balance GPP with respiration (P=R) Attributes used are
Autogenic and Allogenic succession: - GPP(P)
If successional changes are determined by internal co-actions, it is - Respiration (R)
autogenic (self generated) succession.
- NCP
If outside forces effect or control the change, it is the allogenic - Standing Biomass (B)
succession.
- P/R , B/R
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PROCESSES OF SUCCESSION: PROCESSES OF SUCCESSION:
1. NUDATION Development of a bare area without any life form. 4. REACTION The living organisms grow, use water and nutrients
from substratum, and in turn influence the environment which is
2. INVASION Successful establishment of one or more species on
modified through Reaction
a bare area through dispersal or migration followed by ecesis
(establishment) The modification result in environment becoming unsuitable for
existing species and favour newer species (Seral Communities),
As growth and reproduction start, these Pioneer species increase
which replace them.
them
in number and form groups or aggregations
5. STABILIZATION The succession ultimately culminates in a more
3. COMPETITION AND COACTION As number of individuals
or less stable community called Climax which is in equilibrium
grows there is competition, both inter-specific and intra-specific,
with the environment
for space, water and nutrition
The climax community is characterized by maximum biomass and
They influence each other in an number of ways known as
symbiotic linkages between organisms, maintained quite
Coaction
efficiently per unit of available energy
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Diversity richness component increases
Diversity evenness component increases
Life cycle increases in length & complexity
During succession average size of organisms of a system
increases
Small size has the advantage of high metabolic rates, in
nutrient - rich early successional stages.
Larger size and complicated life history is considered as an
adaptation for exploiting seasonal and periodic release of
nutrients and other resources.
Mutualistic symbiosis increases
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Desert Areas
Desert Ecosystem
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Climate Location
The desert is very dry and warm. A very The desert is located 30 degrees
little amount of rain rains a year. Some latitude. Some deserts are in the
deserts can be over 100 degrees F. Thats Mountains. The Mojave desert is the name
so hot
hot. of the desert that is located here in
California.
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Wetlands Wetlands
Wetlands are areas of standing water that support aquatic
plants
Marshes, swamps, and bogs are all considered highest species diversity of all ecosystems
wetlands many species of amphibians, reptiles, birds (such
Plants as ducks and waders), and furbearers can be
adapted to the very moist and humid conditions are called hydrophytes
Pond lilies Cattails Sedges
found in the wetlands
nott considered
id d under
d ffreshwater
h t ecosystems t as
Tamarack Black Spruce there are some, such as salt marshes, that have
high salt concentrationsthese support different
Gum Cypress species of animals, such as shrimp, shellfish, and
various grasses
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River Otter
Wetlands Aquatic Ecosystems
Damselfly Dragonfly Mayfly
Oceans
Marine largest of all the ecosystems
cover about three-fourths of the Earths dominate the Earths surface
surface and include oceans, coral reefs, great diversity of species
and estuaries richest diversity of species even though it contains
fewer species than there are on land
algae
g supply
pp y much of the worlds oxygen
yg
supply and take in a huge amount of
atmospheric carbon dioxide
evaporation of the seawater provides
rainwater for the land
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Intertidal Zone
where the ocean meets the land
sometimes submerged and at other times
exposed
waves and tides come in and out
communities are constantly changing
Oceans
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Mesopelagic Zone
ctenophore related to jellyfish
"twilight zone" of the ocean Big Scale - ambush predator cilia can be illuminated
photic zone above
darkness below
food becomes scarce some animals
migrate up to the surface at night to feed
rely on food that falls down from above Fi fl squid
Firefly id
three kinds of photophore
eat each other Hatchet Fish
sometimes the only things to eat may be bigger than the hunter only a few inches long Viperfish
developed long sharp teeth, specially adapted hinged skull
expandable jaws and stomachs Dragonfish - stomachs hold big meals
Snipeel
up to 1.2m Siphonophores are colonies of animals
related to jellyfish
best known is Portugese Man of War
http://oceanlink.island.net/oinfo/deepsea/meso.html
Narcomedusa
Vampire Squid
Bathypelagic Zone Snake Dragon
Angler Fish
extends down from 1000 to 4000m
only light is from bioluminescent organisms
only food is what trickles down from above, or from eating
Amphi - crustacean
other animals
water
t pressure att this
thi depth
d th is
i considerable
id bl (~100
( 100 400
atmospheres) Ctenophore voracious predator
most animals are either black or red in color Deepstaria very slow swimmers,
very little blue/green light penetrates this deep red is not no tentacles, close flexible bells
reflected and looks black (up to a meter across) around
their prey
Big Red
grows to over
a meter across
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Thank You!
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