Documenti di Didattica
Documenti di Professioni
Documenti di Cultura
SYSTEMATIC DESCRIPTIONS
Order CONODONTOPHORIDA
Eichenberg, 1930
o Acanthodus
Microscopic toothlike and platelike struc- Scolopodus
tures belonging to an unknown monophyle- Acontiodus
tic group of extinct marine animals which ° Belodus
probably were bilaterally symmetrical, soft- Oistodus
bodied, and pelagic. These structures, com- Scandodus
monly called conodonts, are usually either Distacodus
grayish-black or some shade of brown, com- Acodus
posed chiefly of calcium phosphate, and Drepanodus
consist of lamellae, open aborally, that were Paltodus
accreted about an initial pit—the apex of the ° Cordylodus
pulp cavity. Separation of the aforemen- o Ptiloconus
tioned lamellae from one another—along Ulrichodina
one or more growth axes and in one or Stereoconus
more directions—resulted in the formation o Microcoelodus
i
of fanglike structures without denticles, o Strachanognathus
fanglike structures with denticles, denti- Mixoconus
culated blades and bars, and platelike struc-
tures with platforms and/or a greatly ex- FIG. 21 A. Stratigraphic distribution of conodonts
panded pulp cavity. The function per- (Hass, n ) . Classification of genera in families is
formed by conodonts is as yet undeter- indicated by presence or absence of symbol (Dista-
mined. Generally accepted range, L.Ord.- codontidae, o—Belodontidae).
U.Trias.; possible range, U.Cam.-U.Cret.
Acodus PANDER, 1856 (p. 21) [*A. erectus; SD
ULRICH & BASSLER, 1926 (p. 7)]. Resembles
UTILITARIAN Distacodus but asymmetrical, having ridge along
CLASSIFICATION mid-line of one lateral side only. L.Ord.-M.SiL,
?U.Sil.-?LMiss., N.Am.-Eu. FIG. 22,8. *A.
erectus, L.Ord.(Glaukonitsand), Bait.; 8a,b, inner,
Family DISTACODONTIDAE Bassler, outer, mag. unknown (56). FIG. 22,8C. A.
1925 acutus PANDER, L.Ord.(Glaukonitsand), Bait.;
[nom. correct. HASS, 1958 {pro Distacodidae BASSLER, 1925)] diagram, horiz. sec, mag. unknown (52).
Pulp cavity surmounted by a single, Acontiodus PANDER, 1856 (p. 28) [*A. latus; SD
straight or curved, undenticulated, fanglike ULRICH & BASSLER, 1926 (p. 7)] [=Acodina
cusp. L.Ord.-USiL, ?Dev., ?Miss. STAUFFER, 1940]. Compressed anteroposteriorly;
posterior side commonly with median- ridge; in
The stratigraphic distribution of genera horizontal section oudine of anterior side convex
included in the Distacodontidae and Belo- to obtuse; posterior side concave. L.Ord.-M.Ord.,
dontidae is shown graphically in Figure ?U.Ord.-?U.Dev., N.Am.-Eu. FIG. 22,9a-c. *A.
21A. latus, L.Ord.(Glaukonitsand), Bait.; 9a-c, post.,
Distacodus HINDE, 1879 (p. 357) [pro Machairodus ant, diagram, horiz. sec, mag. unknown (52).
PANDER, 1856 {non KAUP, 1833)] [*Machairodus Drepanodus PANDER, 1856 (p. 20) [non MENGE,
incurvus PANDER, 1856; SD MILLER, 1889 (p. 1869] [*D. arcuatus; SD MILLER, 1889 (p. 313)]
313)J [—Machairodia SMITH, 1907]. Bilaterally [=Oneotodus LINDSTROM, 1954]. Almost bilat-
symmetrical; anterior and posterior sides sharp- erally symmetrical; outline biconvex to subcircular
edged; lateral sides with ridge along mid-line. in horizontal section; anterior and posterior sides
L.Ord.-M.SiL, ?U.Sil.-?L.Miss., N.Am.-Eu. rounded or sharp-edged. L.Ord.-U.Sil., ?L.Dev.-
FIG. 22,la,b. *D. incurvus (PANDER); both lat., ?L.Miss., N.Am.-S.Am.-Eu.-Austral. FIG. 22,
mag. unknown. FIG. 22,1C. D. ensiformis 10. D. subarcuatus FURNISH, L.Ord.(Shakopee
(PANDER), L.Ord.(Glaukonitsand), Bait.; diagram, Dol.), USA(Wis.); 10a,b, lat., lat. with diagram,
horiz. sees., mag. unknown (52). horiz. sec, X25 (30).
Miscellanea—Con odonts
Mixoconus SWEET, 1955 (p. 244) [*M. primus]. sides truncated, rounded, grooved, or sharp-edged;
Resembles Distacodus; bilaterally symmetrical or lateral sides commonly costate. L.Ord.-U.Sil.t ?L.
nearly so; anterior and posterior sides rounded; Dev.-?LMiss., N.Am.-Eu.-Austral. FIG. 22,
lateral sides with broadly rounded, aborally wid- lla-c. *P. subaequalis, L.Ord.(Glaukonitsand),
ening ridge; pulp cavity shallow. M.Ord., N.Am. Bait.; lla-c, ant., lat., lat., mag. unknown (52).
FIG. 22,5a,b. *M. primus, Harding Ss., USA FIG. 22,1 Id. P. truncatus PANDER; diagram.
(Colo.), both lat., X27 (72). horiz. sec, mag. unknown (52).
Oistodus PANDER, 1856 (p. 27) [*O. lanceolatus; Scandodus LINDSTROM, 1954 (p. 592) [*S. fur-
SD ULRICH & BASSLER, 1926 (p. 7 ) ] . Like Dista- nishi]. Asymmetrical with anterior and posterior
codus but with base greatly expanded posteriorly. sides sharp-edged. Base expanded on inner side.
Ord., N.Am.-S.Am.-Eu.-Austral. FIG. 22,4a-c. Carina may be present along mid-line of lateral
*O. lanceolatus, L.Ord.(Glaukonitsand), Bait.; sides. Ord., N.Am.-Eu.-Austral. FIG. 22,7. *S.
4a-c, lat., lat., aboral, mag. unknown (52). jurnishi, L.Ord.(L. Planilimbata Z.), Swed.; inner
FIG. 22,4d. O. acuminatus PANDER; diagram, lat., X20 (44).
horiz. sec. above base, mag. unknown (52). Scolopodus PANDER, 1856 (p. 25) [*S. sublaevis;
Paltodus PANDER, 1856 (p. 24) [*P. subaequalis; SD ULRICH & BASSLER, 1926 (p. 7 ) ] . Anterior
SD ULRICH & BASSLER, 1926 (p. 7)] [=Pandero- side rounded to sharp-edged; posterior and lateral
dus ETHINGTON, 1959]. Anterior and posterior sides costate, grooved. L.Ord.-M.Ord.j ?U.Ord.-
Systematic Descriptions
?U.Dev., N.Am.-Eu. FIG. 22,5. *S. sublaevis,
L.Ord.(Glaukonitsand), Bait.; 3a,b, diagram,
horiz. sec, mag. unknown (52).
Stereoconus BRANSON & MEHL, 1933 (p. 27) [*S.
gracilis]. Bilaterally symmetrical, broadest near
rounded posterior side; aboral side with cordate
outline; pulp cavity at notched posterior end. L.
Ord.-M.Ord., N.Am.-Eu. FIG. 22,6. *S.
gracilis, M.Ord.(Harding Ss.), USA(Colo.); lat.,
X25 (7).
Ulrichodina FURNISH, 1938 (p. 334) [*U. prima).
Bilaterally symmetrical; broadest near rounded
anterior side; posterior side sharp-edged; base
indented anteriorly. L.Ord., N.Am. FIG. 22,2.
*U. prima, Shakopee Dol., USA(Wis.); 2a,b, ant.,
lat., X45; 2c, diagram, lat. and horiz. sec. above
base, X25 (30).
Microcoelodus BRANSON & MEHL, 1933 (p. 89) Main cusp indistinct, not terminal; an-
[*M. typus]. Lateral sides of expanded base denti- terior bar or blade short. M.Ord.-L.Miss.
culated. M.Ord., N.Am. FIG. 23,2. *M. typus (L. Osag.).
Joachim Dol., USA(Mo.); post, lat., X25 (8). Coleodus BRANSON & MEHL, 1953 (p. 24) [*C.
Ptiloconus SWEET, 1955 (p. 245) [pro Pteroconus simplex]. Bladelike; denticles closely set; aboral
BRANSON & MEHL, 1933 (non HINDE in Fox, side grooved posterior to pulp cavity and sharp-
1900)] [*Pteroconus gracilis BRANSON & MEHL, edged anterior to pulp cavity. M.Ord., N.Am.
Miscellanea—Conodonts
Trias.
Perm.
Miss.
1
Ord.
Dev.
Sil.
lat., X15 (7).
Arcugnathus COOPER, 1943, in COOPER & SLOSS (p.
Loxodus
172) [*A. tenuis]. COOPER'S description; "Bar
Cyrtoniodus
slender, regularly bowed upward; anterior end
Pachysomia denticulated similar to Hindeodella with alter-
Subprioniodus nating upright teeth; posterior denticles strongly
Paracordylodus i
inclined backward; bar terminating in long slender
Gothodus • denticle; no main cusp present." U.Dev., N.Am.
Tetraprioniodus FIG. 24,J. *A. tenuis, Can.(Alta.); lat, X30
Trichonodella H I ••H i (84).
Leptochirognathus i Bactrognathus BRANSON & MEHL, 1941 (p. 98) [*B.
Loxognathus hamata]. Posterior bar straight; anterior bar flexed
Phragmodus inward. Denticles closely set. Lateral expansions
Plectodina of pulp cavity variform. L.Miss.(uppermost Kin-
Zygognathus derhoo\.-lowermost Osag.), N.Am. FIG. 24,7.
Peri don • B. penehamata HASS, Chappel Ls., USA(Tex.);
Trucherognathus 1
7a-c, oral, aboral, outer lat., X20 (38).
Branmehla HASS, 1959 (p. 381) [*Spathodus in-
Curtognathus •
ornatus BRANSON & MEHL, 1934]. Bladelike; an-
Erismodus •
terior end may be flexed inward slightly; denticles
Roundya
closely set; aboral side sharp-edged; lips of pulp
Chirognathus 1
cavity generally prominent. U.Dev.-LMiss.; N.
Hibbardella Am.-Eu.-Afr. FIG. 24,i. *B. inornata (BRAN-
Ligonodina SON & MEHL), U.Dev.(Houy F.), USA(Tex.);
Neoprioniodus inner lat., X20 (38).
Pravognathus 1 Hindeodina HASS, 1959 (p. 382) [*H. simplaria].
Coleodus t Like Hindeodella but with aborted main cusp;
Rhynchognathodus 1 aboral side sharp-edged; lips of pulp cavity ex-
Holodontus 1 tremely small or entirely absent. U.Dev.-U.Miss.,
Keislognathus , N.Am.-Eu. FIG. 24,5. *H. simplaria, L.Miss.
Rhipidognathus (Chappel Ls.), USA (Tex.); inner lat., X25 (38).
Hindeodel In
1 I I I IVJWwvJv • • >-*
Pravognathus STAUFFER, 1936 (p. 79) [pro Hetero-
gnathus STAUFFER, 1935 (non GIRARD, 1854; nee
L.L/1 1C11IJU 1 n Q wmm
SCHMARDA, 1859; nee KING, 1864; nee REY,
Synprioniodina • • •nun
1888) J [*Heterognathus idonea STAUFFER,'1935].
Cervicornoides
Bladelike, largest denticles above pulp cavity;
Euprioniodina tm iiiiniii IIIIIIIII IIIIIIIII ill
aboral side grooved, broadly so at anterior end.
Hindeodel loides m M.Ord., N.Am. FIG. 24,2. *P. idonea (STAUF-
Hindeodina m• • FER), Decorah Sh., USA(Minn.); inner lat., X35
Apatognathus (68).
Metaprioniodus Trucherognathus BRANSON & MEHL, 1933 (p. 84)
Diplododella [*T. distorta]. Anterior and posterior bars aligned;
Branmehla denticles irregular; those of mature specimen
Avignathus may be located along inner side of massive plat-
Scutula form. M.Ord., N.Am. FIG. 24,4. *T. distorta
Elsonella Joachim Dol., USA(Mo.); outer lat., X25 (8).
Tripodellus
Subfamily HINDEODELLINAE Hass, 1959
Arcugnathus
Bactrognathus Main cusp distinct, not terminal; anterior
Kladognathus i barvor blade short. ?L.Sil.t M.Sil.-U.Trias.
Parachirognathus Hindeodella BASSLER, 1925 (p. 219) [*H. subtilis
ULRICH & BASSLER, 1926; SD (p. 3 8 ) ] . Bar
denticles closely set, commonly with group of
smaller denticles alternating with larger ones;
FIG. 23 A. Stratigraphic distribution of conodonts main cusp generally much larger than bar den-
(Hass, n ) . Classification of genera assigned to fam- ticles; pulp cavity small. 9L.SH., M.Sil.-U.Trias.;
ily Coleodontidae. N.Am.-Eu.-Afr. FIG. 25,2. *H. subtilis, U.Dev.
(Chattanooga Sh.), USA(Tenn.); inner lat., X20
(75).
Systematic Descriptions W47
Cervicornoides STAUFFER, 1938 (p. 424) [*C.
alternatus]. Resembles Hindeodella but is more
sinuous; base of main cusp expanded on inner
side; pulp cavity prominent. M.Dev.-U.Dev.; N.
Am.-Eu. FIG. 25,5. *C. alternatus, U.Dev.
(Olentangy Sh.), USA (Ohio); inner, X50 (69).
Kladognathus REXROAD, 1958 (p. 19) [pro Clado-
gnathus REXROAD, 1957 (non BURMEISTER, 1847)]
[*Cladognathus prima REXROAD, 1957]. Anterior
bar aligned with posterior bar; inner lateral process
directed downward and backward. UMiss.
(Chester.); N.Am. FIG. 25,5. *K. prima, USA
(111.); Sa,b, inner lat., aboral, X40 (55).
Metaprioniodus HUDDLE, 1934 (p. 57) [*M. bi-
angulatus]. Resembles Hindeodella but has dis-
crete denticles with largest denticles near pos-
terior deflection. V' .Dev.-LMiss.(Kinderhoo\.);
N.Am. FIG. 25,4. *M. biangulatus, L.Miss.
(New Albany Sh.), USA(Ind.); 4a,b, inner lat.,
outer lat., X15 (42).
Tripodellus SANNEMANN, 1955 (p. 155) [*T.
flexuosus]. Posterior, anterior, and inner lateral
bars compressed, denticulated; anterior bar much
larger than posterior and inner lateral bars,
directed downward, and oriented so as to face
slightly toward outer side of unit; inner lateral
bar joined to front basal portion of main cusp,
curved backward and directed downward slightly;
main cusp with sharp-edged anterior and pos-
terior sides, curved inward and backward slightly;
Bactrognathus aboral side sharp-edged; pulp cavity very small.
FIG. 24. Coleodontidae (Coleodontinae) (p. W45- U.Dev., Eu. FIG. 25,2. *T. flexuosus, L. Chei-
W46). loceras Z., Eu.; outer lat., X40 (61).
Cervicornoides
Metaprioniodus
INDEX
Names included in the following index are classified typographically as follows: (1)
Roman capital letters are used for suprafamilial taxonomic units which are recognized as
valid in classification; (2) italic capital letters are employed for suprafamilial categories
which are considered to be junior synonyms of valid names; (3) morphological terms and
generic family names accepted as valid are printed in roman type; and (4) generic and
family names classed as invalid, including junior homonyms and synonyms, are printed in
italics. Page numbers printed in boldface type (as W134) indicate the location of systematic
descriptions or definitions of morphological terms.
ABEL, W179, Wl80 Annelidichnium, Wl83 Arenicolithes, Wl89
Abeliella, W228 A1lnulati, WI07, WI13 Arenieolitidae, Wl81
aboral, WS Annulosi, WI07, WllO Arenicoloides, Wl89
aboral attachment scar, WS Anobichnium, W228 Aristophycus, W232
aboral cavity, WS Anomalophycus, W226 Armelia, W223
aboral edge, WS anterior, W6 Arthraria, WI84
aboral extension, WS anterior arch, W6 Arthriehnites, Wl84
aboral groove, WS anterior bar, W6 Arthrodendron, W224
aboral margin, WS anterior blade, W6 Arthrophicus, Wl84
aboral process, W5 anterior curvature, W6 Arthrophycus, WI84
aboral projection, WS anterior deflection, W6 Artiodaetylus, W201
aboral side, WS, Wl7 anterior denticles, W6 Asabellarifex, WI84
aboral surface, W6 anterior edge, W6 Asaphoidichnus, WI84
aboro-lateral groove, W6 anterior face, W6 ASCARIDATINA, WI46, W148
Acanthichnus, WI83 anterior inner bar, W6 Asehemonia, W2I8
Acanthodus, W4S anterior inner lateral bar, W6 a-side, W5
Acanthus, W238 anterior limb, W6 Aspidella, W232
accessory lobes, W6 anterior margin, W6 Aspidiaria, W234
Aeodina, W43 anterior outer bar, W6 assemblage, W7
Acodus, W39, W43 anterior outer lateral bar, W6 Astaeoderma, W65
Acontiodus, W43 anterior process, W6 Asteriacites, WI84
Acripes, Wl91 anterior projection, W6 Asterichnites, WI84
Aenigmichnus, W232 anterior side, W6 Asterophycus, WI84
Aeolisaccus, W223 anterior wing, W6 Asterosalpinx, WIS6
Aequorfossa, W238 antero-inferior process, W6 Asterosoma, WI84
Agarites, W238 antero-inner-lateral flange, W6 Astrophora, W206
Aglaopheniolites, WI83 Antholithina, W232 Astropolithon, W184
Alcyonidiopsis, Wl94 anticusp, W6 Atikokania, W232
Aleetoruridae, Wl80 Anzalia, W223 Atollites, WI84
Aleetorurus, W220 Apatognathus, WS3 attachment scar, W7
Algaeites, W179, Wl87 apex, W6 Aulaeomerella, W139
Algites, W220 Aphelognathus, WS6 Aulacophyeus, W208
Amanlisia, W238 Aphrodita, WISI Aulichnites, WI85
Amansites, W238 Aphroditidae, W146, WI 51 AUSTIN, WIOO
Ambalodus, WS8 Aphrostoma, W226 Avignathus, WSO
Ambrolinevitus, WI27 apical cone, W7 axis, WIO
Amiskwia, W148 apical denticle, W7 azygous node, WIO
Amiskwidae, W148 apical lamella, W7
Amiskwiidae, W146, W148 apical lip, W7 Bactrognathus, W40, W46
Amorphognathus, WS8 apical pit, W7 Bactrotheca, WI2S
Ampelichnus,W238 Apodichnites, W220 Bactryllium, W223
Amphictene, WI63 appressed denticles, W7 Baieropsis, W236
Amphictenidae, W146, WI63 apron, W7 Balanoglossites, WI8S
Amphinome, WI53 Arabellites, W149 Balognathinae, W42, W62
Amphinomidae, W146, WI53 arch, W7 Balognathus, W39, W62
Ancyrodella, W32, W40, WS8, Archaeonassa, W183 Banffia, WI69
wn, W89 Archaeophyton, W232 bar, WIO
Ancyrodelloides, WS8 Archaeorrhiza, W238 Barella, W132
Ancyrognathus, W40, W58, W83, Archaeoscolex, W238 BARRANDE, WIOO, WI07, Wl16
W86 Archaeozoon, W226 bar teeth, WIO
Aneyroides, W58, W248 Archarenicola, WI63 basal attachment scar, WIO
Aneyrolepis, W248 Areheognathus, W65 basal canalules, WIO
Aneyropenta, W58 Arcugnathus, W46 basal cavity, WIO, Wl7
Angulodus, WSS Arenicola, WI63 basal cavity inverted, WIO
ANNELIDA, W145, W146, Arenicolidae, W146, WI63 basal cone, WIO, Wl8
W148 Arenicolites, W182, WI83 basal excavation, WIO