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TypesofStomatainPlants|PlantScience4U

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Stomata are minute pores which occur on epidermal surface of leaves and also some herbaceous
stems.Eachstomaisguardedbytwospecialisedepidermalcells,calledguardcells.Theseguardcells
are also surrounded by other specialised epidermal cells called subsidiary cells or accessory cells.
Thesecellsalsoplaysanimportantroleduringopeningandclosingofstomata.

Metcalfe and Chalk (1950) classified stomata on the basis of number and arrangement of the
subsidiarycellsintothefollowingtypes:

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1.Anomocytic(irregularcelled)orRanunculaceous:Inthistype,thestomataremainssurrounded

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TypesofStomatainPlants|PlantScience4U

bylimitednumberofsubsidiarycellswhicharequitealiketheremainingepidermalcells.
Example:Ranunculaceae,Malvaceae,Papaveraceae
2.Anisocytic(Unequalcelled)orCruciferous:Inthisstomataremainssurroundedbythree
subsidiarycellsofwhichoneisdistinctlysmallerthantheothertwo.

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Epigealvs
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Example:Cruciferacea,Solanum,Nicotianaetc.
3.Paracytic(Parallelcelled)orRubiaceous:Inthistype,thestomatasurroundedbytwosubsidiary
cellswhichareparalleltothelongitudinalaxisofporeandguardcells.

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Evolutionin
Pteridophytes

4.Diacytic(Crosscelled)orCaryophyllaceous:Inthistype,thestomataremainssurroundedbya
pairofsubsidiarycellswhosecommonwallisatrightanglestotheguardcells.
Example:Acanthacea,Caryophyllaceae
5.Actinocytic:Thesestomataaresurroundedbyfourormoresubsidiarycells,elongatedradiallyto
thestomata.

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Example:Araceae,Musaceae,Commelinaceae
6.Cyclocytic:Thestomataaresurroundedbyfourormoresubsidiarycellsarrangedinanarrowring

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aroundthestoma
Example:Palmae,Pandanus,Cyclanthaceae

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7.Graminaceoustype:Thestomatalguardcellsaredumbbellshaped.Theyaresurroundedby
Name

subsidiarycellswhicharelyingparalleltothelongaxisofthepore.
Example:InthemembersofPoaceaeandcyperaceae

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Labels:Actinocyticstomata,Anisocyticstomata,Anomocyticstomata,classificationofstomata,
Cyclocyticstomata,Diacyticstomata,Graminaceoustype,MetcalfeandChalk,Paracyticstomata,
stomata,StomataTypes

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